Abirchis a thin-leaveddeciduoushardwoodtreeof thegenusBetula(/ˈbɛtjʊlə/),[2]in the familyBetulaceae,which also includesalders,hazels,andhornbeams.It is closely related to thebeech-oakfamilyFagaceae.The genusBetulacontains 30 to 60known taxaof which 11 are on theIUCN 2011 Red Listof Threatened Species. They are typically short-livedpioneer speciesand are widespread in theNorthern Hemisphere,particularly in northern areas oftemperateclimates and inboreal climates.[3]Birch woodis used for a wide range of purposes.
Birch Temporal range:
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Betula pendula(Silver birch) | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Rosids |
Order: | Fagales |
Family: | Betulaceae |
Subfamily: | Betuloideae |
Genus: | Betula L. |
Subgenera | |
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Range ofBetula | |
Synonyms[1] | |
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Description
editThis sectionneeds additional citations forverification.(March 2023) |
Birch species are generally small to medium-sizedtreesorshrubs,mostly of northerntemperateand boreal climates.[4]The simpleleavesare alternate, singly or doubly serrate, feather-veined, petiolate and stipulate. They often appear in pairs, but these pairs are really borne on spur-like, two-leaved, lateralbranchlets.[5]The fruit is a smallsamara,although the wings may be obscure in some species. They differ from thealders(Alnus,another genus in the family) in that the femalecatkinsare not woody and disintegrate at maturity, falling apart to release the seeds, unlike the woody, cone-like female alder catkins.
Thebarkof all birches is characteristically marked with long, horizontallenticels,and often separates into thin, papery plates, especially upon thepaper birch.[6]Distinctive colors give the common namesgray,white,black,silverandyellowbirch to different species.[7]
The buds, forming early and full-grown by midsummer, are all lateral, without a terminal bud forming; the branch is prolonged by the upper lateral bud. The wood of all the species is close-grained with a satiny texture and capable of taking a fine polish; its fuel value is fair.
Flower and fruit
editThe flowers aremonoecious,and open with or before the leaves. Once fully grown, these leaves[clarification needed]are usually3–6 millimetres (1⁄8–1⁄4in) long on three-flowered clusters in the axils of the scales of drooping or erectcatkinsor aments. Staminate catkins are pendulous, clustered, or solitary in the axils of the last leaves of the branch of the year or near the ends of the short lateral branchlets of the year. They form in early autumn and remain rigid during the winter. The scales of the mature staminate catkins are broadly ovate, rounded, yellow or orange colour below the middle and dark chestnut brown at apex. Each scale bears two bractlets and three sterile flowers, each flower consisting of a sessile, membranous, usually two-lobed, calyx. Each calyx bears four short filaments with one-celled anthers or strictly, two filaments divided into two branches, each bearing a half-anther. Anther cells open longitudinally. The pistillate segments are erect or pendulous, and solitary, terminal on the two-leaved lateral spur-like branchlets of the year. The pistillate scales are oblong-ovate, three-lobed, pale yellow-green often tinged with red, becoming brown at maturity. These scales bear two or three fertile flowers, each flower consisting of a naked ovary. The ovary is compressed, two-celled, and crowned with two slender styles; the ovule is solitary. Each scale bears a single small, winged nut that is oval, with two persistent stigmas at the apex.
Taxonomy
editThis sectionneeds additional citations forverification.(April 2021) |
Subdivision
editBetulaspecies are organised into five subgenera.
- Birches native toEurasiainclude
- Betula albosinensis– Chinese red birch (northern + central China)
- Betula alnoides– alder-leaf birch (China, Himalayas, northern Indochina)
- Betula ashburneri– (Bhutan, Tibet, Sichuan, Yunnan Provinces in China)
- Betula baschkirica– (eastern European Russia)
- Betula bomiensis– (Tibet)
- Betula browicziana– (Turkey and Georgia)
- Betula buggsii– (China)
- Betula calcicola– (Sichuan + Yunnan Provinces in China)
- Betula celtiberica– (Spain)
- Betula chichibuensis– (Chichiburegion of Japan)[8]
- Betula chinensis– Chinese dwarf birch (China, Korea)
- Betula coriaceifolia– (Uzbekistan)
- Betula corylifolia– (Honshu Island in Japan)
- Betula costata– (northeastern China, Korea, Primorye region of Russia)
- Betula cylindrostachya– (Himalayas, southern China, Myanmar)
- Betula dahurica– (eastern Siberia, Russian Far East, northeastern China, Mongolia, Korea, Japan)
- Betula delavayi– (Tibet, southern China)
- Betula ermanii– Erman's birch (eastern Siberia, Russian Far East, northeastern China, Korea, Japan)
- Betula falcata– (Tajikistan)
- Betula fargesii– (Chongqing + Hubei Provinces in China)
- Betula fruticosa– (eastern Siberia, Russian Far East, northeastern China, Mongolia, Korea, Japan)
- Betula globispica– (Honshu Island in Japan)
- Betula gmelinii– (Siberia, Mongolia, northeastern China, Korea, Hokkaido Island in Japan)
- Betula grossa– Japanese cherry birch (Japan)
- Betula gynoterminalis– (Yunnan Province in China)
- Betula honanensis– (Henan Province in China)
- Betula humilisorBetula kamtschatica– Kamchatka birchplatyphylla(northern + central Europe, Siberia, Kazakhstan, Xinjiang, Mongolia, Korea)
- Betula insignis– (southern China)
- Betula karagandensis– (Kazakhstan)
- Betula klokovii– (Ukraine)
- Betula kotulae– (Ukraine)
- Betula luminifera– (China)
- Betula maximowicziana– monarch birch (Japan, Kuril Islands)
- Betula medwediewii– Caucasian birch (Turkey, Iran, Caucasus)
- Betula megrelica– (Republic of Georgia)
- Betula microphylla– (Siberia, Mongolia, Xinjiang, Kazakhstan, Kyrgyzstan, Uzbekistan)
- Betula nana– dwarf birch (northern + central Europe, Russia, Siberia, Greenland, Northwest Territories of Canada))
- Betula pendula– silver birch (widespread in Europe and northern Asia; Morocco; naturalized in New Zealand and scattered locations in US + Canada)
- Betula platyphylla– (Betula pendulavar.platyphylla)—Siberian silver birch (Siberia, Russian Far East, Manchuria, Korea, Japan, Alaska, western Canada)
- Betula potamophila– (Tajikistan)
- Betula potaninii– (southern China)
- Betula psammophila– (Kazakhstan)
- Betula pubescens– downy birch, also known as white, European white or hairy birch (Europe, Siberia, Greenland, Newfoundland; naturalized in scattered locations in US)
- Betula raddeana– (Caucasus)
- Betula saksarensis– (Khakassiya region of Siberia)
- Betula saviczii– (Kazakhstan)
- Betula schmidtii– (northeastern China, Korea, Japan, Primorye region of Russia)
- Betula sunanensis– (Gansu Province of China)
- Betula szechuanica– (Betula pendulavar.szechuanica)—Sichuan birch (Tibet, southern China)
- Betula tianshanica– (Kazakhstan, Kyrgyzstan, Tajikistan, Uzbekistan, Xinjiang, Mongolia)
- Betula utilis– Himalayan birch (Afghanistan, Central Asia, China, China, Tibet, Himalayas)
- Betula wuyiensis– (Fujian Province of China)
- Betula zinserlingii– (Kyrgyzstan)
Note: many American texts haveB. pendulaandB. pubescensconfused, though they are distinct species with different chromosome numbers.
- Birches native toNorth Americainclude
- Betula alleghaniensis– yellow birch (B. lutea) (eastern Canada, Great Lakes, upper eastern US, Appalachians)
- Betula caerulea– blue birch (northeast of North America)
- Betula cordifolia– mountain paper birch (eastern Canada, Great Lakes, New England US)
- Betula glandulosa– American dwarf birch (Siberia, Mongolia, Russian Far East, Alaska, Canada, Greenland, mountains of western US and New England, Adirondacks)
- Betula kenaica– Kenai birch ( Alaska, northwestern North America)
- Betula lenta– sweet birch, cherry birch, or black birch (Quebec, Ontario, eastern US)
- Betula michauxii– Newfoundland dwarf birch (Newfoundland, Labrador, Quebec, Nova Scotia)
- Betula minor– dwarf white birch (eastern Canada, mountains of northern New England and Adirondacks)
- Betula murrayana– Murray's birch (Great Lakes endemic)
- Betula nana– dwarf birch or bog birch (also in northern Europe and Asia)
- Betula neoalaskana– Alaska paper birch also known as Alaska birch or Resin birch (Alaska and northern Canada)
- Betula nigra– river birch or black birch (eastern US)
- Betula occidentalis– water birch or red birch (B. fontinalis) (Alaska, Yukon, Northwest Territories, western Canada, western US)
- Betula papyrifera– paper birch, canoe birch or American white birch (Alaska, most of Canada, northern US)
- Betula populifolia– gray birch (eastern Canada, northeastern US)
- Betula pumila– swamp birch (Alaska, Canada, northern US)
- Betula uber– Virginia round-leaf birch (southwestern Virginia)
Etymology
editThe common namebirchcomes fromOld Englishbirce,bierce,fromProto-Germanic*berk-jōn(cf.GermanBirke,West Frisianbjirk), an adjectival formation from *berkōn(cf.Dutchberk,Low GermanBark,Danishbirk,Norwegianbjørk), itself from theProto-Indo-Europeanroot *bʰerHǵ-~bʰrHǵ-, which also gaveLithuanianbéržas,LatvianBērzs,Russianберёза (berëza),Ukrainianбереза (beréza),Albanianbredh'fir',Ossetianbærz(æ),Sanskritbhurja,Polishbrzoza,Latinfraxinus'ash (tree)'. This root is presumably derived from *bʰreh₁ǵ-'to shine, whiten', in reference to the birch's white bark. TheProto-Germanicruneberkananis named after the birch.
The generic nameBetulais fromLatin,which is adiminutiveborrowed fromGaulishbetua(cf.Old Irishbethe,Welshbedw).
Evolutionary history
editWithin Betulaceae, birches are most closely related to alder. The oldest known birch fossils are those ofBetula leopoldaefrom theKlondike Mountain Formationin Washington State, US, which date to the earlyEocene(Ypresian) around 49 million years ago.[9]
Ecology
editBirches often form even-aged stands on light, well-drained, particularlyacidic soils.They are regarded aspioneer species,rapidly colonizing open ground especially insecondary successionalsequences following a disturbance or fire. Birches are early tree species to become established inprimary successions,and can become a threat to heathland if theseedlingsand saplings are not suppressed by grazing or periodic burning. Birches are generally lowland species, but some species, such asBetula nana,have amontanedistribution. In theBritish Isles,there is some difference between the environments ofBetula pendulaandBetula pubescens,and some hybridization, though both are "opportunists in steady-state woodland systems".Mycorrhizal fungi,including sheathing (ecto)mycorrhizas, are found in some cases to be beneficial to tree growth.[10]
A large number of lepidopteran insects feed on birch foliage.
Uses
editBecause of the hardness of birch, it is easier to shape it with power tools; it is quite difficult to work it with hand tools.[11]
- Birch wood is fine-grained and pale in colour, often with an attractivesatin-like sheen. Ripple figuring may occur, increasing the value of the timber forveneerand furniture-making. The highly decorative Masur (orKarelian) birch, fromBetula verrucosavar.carelica,has ripple textures combined with attractive dark streaks and lines.[citation needed]
- Birchplywoodis made from laminations of birch veneer. It is light but strong, and has many other good properties. It is among the strongest and dimensionally most stable plywoods, although it is unsuitable for exterior use. Birch plywood is used to makelongboards(skateboard), giving it a strong yet flexible ride. It is also used (often in very thin grades with many laminations) for makingmodel aircraft.[citation needed]
- Extracts of birch are used forflavoringorleather oil,and in cosmetics such assoaporshampoo.In the past, commercial oil of wintergreen (methyl salicylate) was made from thesweet birch(Betula lenta).[citation needed][12]
- Birch-taror Russian oil extracted from birch bark isthermoplasticand waterproof; it was used as aglueon, for example,arrows,and also for medicinal purposes.[13]
- Fragrant twigs of wintergreen group birches are used insaunas.[citation needed]
- Birch is also associated with the feast ofPentecostin Central and Eastern Europe and Siberia, where its branches are used as decoration for churches and homes on this day.[citation needed]
- Ground birch bark,fermentedinsea water,is used for seasoning the woolen, hemp or linen sails and hemp rope oftraditional Norwegian boats.[citation needed]
- Birch twigs bound in a bundle, also called birch, were used forbirching,a form ofcorporal punishment.[citation needed]
- ManyNative Americans in the United StatesandIndigenous peoples in Canadaprize the birch for its bark, which because of its light weight, flexibility, and the ease with which it can be stripped from fallen trees, is often used for the construction of strong, waterproof but lightweightcanoes,bowls, andwigwams.[citation needed]
- TheHughes H-4 Herculeswas made mostly of birch wood, despite its better-known moniker, "The Spruce Goose".
- Birch plywood was specified by the BBC as the only wood that can be used in making the cabinets of the long-livedLS3/5Aloudspeaker.[14]
- Birch is used asfirewoodbecause of its highcalorific valueper unit weight and unit volume. It burns well, without popping, even when frozen and freshly hewn. The bark will burn very well even when wet because of the oils it contains. With care, it can be split into very thin sheets that will ignite from even the smallest of sparks. Birch wood can be used to smoke foods.[citation needed]
- Birch seeds are used asleaf litterin miniature terrain models.[15]
- Birch oil is used in the manufacture ofRussia leather,a water-resistant leather.
As food
editThe inner bark is considered edible as an emergency food, even when raw.[12]It can be dried and ground into flour, as was done by Native Americans and early settlers. It can also be cut into strips and cooked like noodles.[12]
Thesapcan be drunk or used to makesyrup[12]andbirch beer.Tea can be made from the red inner bark of black birches.[12]
Cultivation
editWhite-barked birches in particular are cultivated as ornamental trees, largely for their appearance in winter. The Himalayan birch,Betula utilis,especially the variety or subspeciesjacquemontii,is among the most widely planted for this purpose. It has been cultivated since the 1870s, and manycultivarsare available, including 'Doorenbos', 'Grayswood Ghost' and 'Silver Shadow'; 'Knightshayes' has a slightly weeping habit. Other species with ornamental white bark includeBetula ermanii,Betula papyrifera,Betula pendulaandBetula raddeana.[16]
Medical
editApproved topical medicine
editIn the European Union, aprescriptiongelcontaining birch bark extract (commercial nameEpisalvan,betulae cortex dry extract (5–10: 1); extraction solvent:n-heptane95% (w/w)) was approved in 2016 for thetopical treatmentof minor skin wounds in adults.[17]Although itsmechanism of actionin helping to heal injured skin is not fully understood, birch bark extract appears to stimulate the growth ofkeratinocyteswhich then fill the wound.[17][18]
Research and traditional medicine
editPreliminary research indicates that thephytochemicals,betulinand possibly othertriterpenes,are active inEpisalvangel and wound healing properties of birch bark.[18]
Over centuries, birch bark was used intraditional medicinepractices byNorth American indigenous peoplefor treating superficial wounds by applying bark directly to the skin.[18]Splintsmade with birch bark were used ascasts for broken limbsin the16th century.[19]
Paper
editWood pulpmade from birch gives relatively long and slender fibres for ahardwood.The thin walls cause the fibre to collapse upon drying, giving apaperwith low bulk and low opacity. The birch fibres are, however, easily fibrillated and give about 75% of the tensile strength of softwood.[clarification needed][20]The low opacity makes it suitable for makingglassine.
InIndia,the birch (Sanskrit:भुर्ज,bhurja) holds great historical significance in the culture ofNorth India,where the thin bark coming off in winter was extensively used as writing paper. Birch paper (Sanskrit: भुर्ज पत्र,bhurja patra) is exceptionally durable and was the material used for many ancient Indian texts.[21][22]TheRomanperiodVindolanda tabletsalso use birch as a material on which to write and birch bark was used widely in ancientRussiaas notepaper (beresta) and for decorative purposes and even makingfootwear(lapti) and baskets.[citation needed]
Use in musical instruments
editBirch wood is sometimes used as atonewoodfor semiacoustic andacoustic guitarbodies, and occasionally for solid-body guitar bodies. It is also a common material used inmalletsforkeyboard percussion.[citation needed]Drum manufacturers, such asGretschandYamaha,have been known to use birch wood in the construction of drum shells, owing to its strength and colour which takes stain in an appealing way, and which can also amber over very well, while also giving the drums an appealing tone which changes depending on the type of birch used.[23]
Culture
editBirches have spiritual importance in several religions, both modern and historical. In Celtic cultures, the birch symbolises growth, renewal, stability, initiation, and adaptability because it is highly adaptive and able to sustain harsh conditions with casual indifference. Proof of this adaptability is seen in its easy and eager ability to repopulate areas damaged by forest fires or clearings. Birches are also associated withTír na nÓg,the land of the dead and theSidhe,in Gaelic folklore, and as such frequently appear in Scottish, Irish, and English folksongs and ballads in association with death, or fairies, or returning from the grave. The leaves of the silver birch tree are used in the festival of St George, held inNovosejand other villages in Albania.[24]
The birch isNew Hampshire's state tree and the national tree of Finland and Russia. Theyellow birchis the official tree of the province of Quebec (Canada). The birch is a very important element in Russian culture and represents the grace, strength, tenderness and natural beauty of Russian women as well as the closeness to nature of the Russians.[25]It's associated with marriage and love.[26]There are numerous folkloric Russian songs in which the birch tree occurs. TheOrnäs birchis the national tree of Sweden. TheCzechword for the month of March, Březen, is derived from the Czech word bříza meaning birch, as birch trees flower in March under local conditions. The silver birch tree is of special importance to the Swedish city ofUmeå.In 1888, theUmeå city firespread all over the city and nearly burnt it down to the ground, but some birches, supposedly, halted the spread of the fire. To protect the city against future fires, wide avenues were created, and these were lined with silver birch trees all over the city. Umeå later adopted the unofficial name of "City of the Birches (Björkarnas stad) ". Also, the ice hockey team of Umeå is calledBjörklöven,translated to English "The Birch Leaves".[citation needed]
"Swinging" birch trees was a common game for American children in the nineteenth century. American poetLucy Larcom's "Swinging on a Birch Tree" celebrates the game.[27]The poem inspiredRobert Frost,who pays homage to the act of climbing birch trees in his more famous poem, "Birches".[28]Frost once told "it was almost sacrilegious climbing a birch tree till it bent, till it gave and swooped to the ground, but that's what boys did in those days".[29]
See also
editReferences
edit- ^"World Checklist of Selected Plant Families: Royal Botanic Gardens, Kew".apps.kew.org.
- ^Sunset Western Garden Book,1995:606–607
- ^Ashburner, K. & McAllister, H.A. (2013). The genusBetula:a taxonomic revision of birches: 1–431. Royal Botanic Gardens. Kew.
- ^"11 Birch Trees Common to North American Landscapes".The Spruce.Retrieved2022-04-27.
- ^Keeler, Harriet L. (1900).Our Native Trees and How to Identify Them.New York: Charles Scribner's Sons. pp.295–297.
- ^"Recognizing Trees Using Bark".Cowling ArboretumCarleton College.Retrieved2022-04-27.
- ^"Trees of the Adirondacks: Yellow Birch |Betula alleghaniensis".Visitor Interpretive CenterPaul Smith's College.Retrieved2022-04-27.
- ^Kinver, Mark (30 September 2015)."UK team germinates critically endangered Japanese birch".BBC News.BBC.Retrieved30 September2015.
- ^Forest, Félix;Savolainen, Vincent;Chase, Mark W.;Lupia, Richard; Bruneau, Anne; Crane, Peter R. (2005-01-01)."Teasing Apart Molecular- Versus Fossil-based Error Estimates when Dating Phylogenetic Trees: A Case Study in the Birch Family (Betulaceae)".Systematic Botany.30(1): 118–133.doi:10.1600/0363644053661850.ISSN0363-6445.S2CID86080433.
- ^Birches.(A Symposium, Royal Botanic Garden, Edinburgh 24–26 September 1982.Proceedings of the Royal Society of Edinburgh,85B, 1–11, 1984.
- ^ "Birch".Wood Magazine.RetrievedDecember 1,2013.
- ^abcdeAngier, Bradford(1974).Field Guide to Edible Wild Plants.Harrisburg, PA: Stackpole Books. p. 34.ISBN0-8117-0616-8.OCLC799792.
- ^"Birch Tar – How to collect it".Archived fromthe originalon February 27, 2008.
- ^Prakel, David (August 1979)."BBC's Home Service",Hi-Fi Answers,pp67–9 (Courtesy link)
- ^Joyce, Daniel."Birch Seed Leaves".reapermini.com. Archived fromthe originalon 2010-12-31.Retrieved2010-04-09.
- ^Bartlett, Paul (2015). "White-barked birches".The Plantsman.New Series.14(3): 146–151.
- ^ab"Episalvan".European Medicines Agency.5 February 2016.Retrieved29 October2020.
- ^abcEbeling, Sandra; Naumann, Katrin; Pollok, Simone; Wardecki, Tina; Vidal-y-Sy, Sabine; Nascimento, Juliana M.; Boerries, Melanie; Schmidt, Gudula; Brandner, Johanna M.; Merfort, Irmgard (2014-01-22). Simon, Michel (ed.)."From a traditional medicinal plant to a rational drug: Understanding the clinically proven wound healing efficacy of birch bark extract".PLOS ONE.9(1): e86147.Bibcode:2014PLoSO...986147E.doi:10.1371/journal.pone.0086147.ISSN1932-6203.PMC3899119.PMID24465925.
- ^William Arthur Clark (January 1, 1937)."History of fracture treatment up to the sixteenth century".The Journal of Bone and Joint Surgery.19(1): 61–62. Archived fromthe originalon December 3, 2013.RetrievedDecember 2,2013.
Another method cited was that of splints made of birch bark soaked in water until quite soft. They were then carefully fitted to the limb and tied with bark thongs. On drying, they became stiff and firm. There is no record of the use of extension, but, nevertheless, very few crippled and deformed Indians were to be seen.
- ^Nanko, Hiroki; Button, Alan; Hillman, Dave (2005).The World of Market Pulp.US: WOMP, LLC. pp. 192–195.ISBN0-615-13013-5.
- ^Sanjukta Gupta, "Lakṣmī Tantra: A Pāñcarātra Text", Brill Archive, 1972,ISBN90-04-03419-6.Snippet:... the text recommends that the bark of the Himalayan birch tree (bhurja-patra) should be used for scribbling mantras...
- ^Amalananda Ghosh, "An Encyclopaedia of Indian Archaeology", BRILL, 1990,ISBN90-04-09264-1.Snippet:... Bhurja-patra, the inner bark on the birch tree grown in the Himalayan region, was a very common writing material...
- ^Cerra, Steve La (2022-01-21)."A Brief Guide to Drum Tonewoods".Yamaha Music – Blog.Retrieved2024-09-03.
- ^"Traditional celebrations in Novosej".RASP. Archived fromthe originalon December 3, 2013.RetrievedAugust 28,2013.
- ^"The Birch: Russia's Tree | News & Info".kurochkaclothing.com.Retrieved2018-09-24.
- ^Weiss, Peg; Kandinsky, Wassily (1995).Kandinsky and Old Russia: the artist as ethnographer and shaman.New Haven: Yale University Press. p.36.ISBN0300056478.OCLC30701876.
- ^Pfileger, Pat."Our Young Folks: Swinging on a Birch-Tree, by Lucy Larcom & Winslow Homer (1867)".Merry Coz.
- ^Fagan, Deirdre J. (2007).Critical Companion to Robert Frost: A Literary Reference to His Life and Work.Infobase Publishing. p. 42.ISBN978-1-4381-0854-4.Retrieved10 November2013.
- ^Parini, Jay (1999).Robert Frost: A Life.New York:Halt. p.22.ISBN0-8050-3181-2.
Sources
edit- Furlow, John J. (1997)."Betula".In Flora of North America Editorial Committee (ed.).Flora of North America North of Mexico (FNA).Vol. 3. New York and Oxford: Oxford University Press – via eFloras.org,Missouri Botanical Garden,St. Louis, MO &Harvard University Herbaria,Cambridge, MA.
- Li, Pei-chun; Skvortsov, Alexei K."Betula".Flora of China.Vol. 4 – via eFloras.org,Missouri Botanical Garden,St. Louis, MO &Harvard University Herbaria,Cambridge, MA.
- Grimshaw, John (2009).New Trees, Recent introductions to cultivation.Kew Publishing,RBG Kew.pp. 163–174.
- Chisholm, Hugh,ed. (1911). .Encyclopædia Britannica.Vol. 3 (11th ed.). Cambridge University Press.
External links
edit- Tree Family BetulaceaeDiagnostic photos of many species,Morton Arboretumspecimens
- Eichhorn, Markus (July 2010)."The Birch Tree".Test Tube.Brady Haranfor theUniversity of Nottingham.