Huntsman spider

Huntsman spiders,members of thefamilySparassidae(formerly Heteropodidae), catch their prey by hunting rather than in webs.They are also calledgiant crab spidersbecause of their size and appearance. Larger species sometimes are referred to aswood spiders,because of their preference for woody places (forests, mine shafts, woodpiles, wooden shacks). In southern Africa the genusPalystesare known asrain spidersorlizard-eating spiders.Commonly, they are confused withbaboon spidersfrom theMygalomorphaeinfraorder,which are not closely related.

Huntsman spiders Temporal range:
Palystes castaneus,showing sparassid pattern of eyes in two rows of four, with the robust build and non-clavatepedipalpsof a female
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Araneae
Infraorder:	Araneomorphae
Family:	Sparassidae Bertkau,1872
Diversity
88 genera,1363 species

More than a thousand Sparassidae species occur in most warm temperate to tropical regions of the world, including much ofAustralia,Africa,Asia,theMediterranean Basin,and theAmericas.

Several species of huntsman spider can use an unusual form of locomotion. Thewheel spider(Carparachne aureoflava) from theNamibuses acartwheelingmotion which gives it its name, whileCebrennus rechenbergiuses ahandspringmotion.

Description

Sparassids are eight-eyedspiders.The eyes appear in two largely forward-facing rows of four on theanterior aspectof theprosoma.Many species grow very large – inLaos,malegiant huntsman spiders(Heteropoda maxima) attain a legspan of 25–30 centimetres (9.8–11.8 in). People unfamiliar with spider taxonomy commonly confuse large species withtarantulas,but huntsman spiders can generally be identified by their legs, which, rather than being jointed vertically relative to the body, are twisted in such a way that in some attitudes the legs extend forward in acrab-like fashion.It is also commonly confused for abrown reclusespider, due to their shared coloring. However, brown recluse venom is significantly dangerous to humans, while that of the huntsman spider is less so.

On their upper surfaces the main colours of huntsman spiders are inconspicuous shades of brown or grey, but many species have undersides more or lessaposematicallymarked in black-and-white.Their legs bear fairly prominent spines, but the rest of their bodies are smoothly furry. They tend to live under rocks, bark and similar shelters, but human encounters are common in sheds, garages and other infrequently-disturbed places. The banded huntsman (Holconia) is large, grey to brown with striped bands on its legs. The badge huntsman (Neosparassus) is larger still, brown and hairy. The tropical or brown huntsman (Heteropoda) is also large and hairy, with mottled brown, white and black markings. The eyesight of these spiders is not as good as that of theSalticidae(jumping spiders). Nevertheless, their vision is quite sufficient to detect approaching humans or other large animals from some distance.

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  Palystes superciliosus,ventral aspect, showingaposematiccoloration, plus typically masculinegracilebuild andclavatepedipalpsarmed with mating spurs
• 
  A huntsman spider consuming a smallbeetle
• 
  A femaleHeteropoda venatoriaconsuming akatydid
• 
  Adult social huntsmanDelena cancerideson the underside of a log in Victoria, Australia
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  Olios argelasius
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  Unidentified species in Ghana

Identification

They can be distinguished from otherspiderfamilies by their appearance, as other spiders similar to them are smaller in size. They are often confused fortarantulasdue to their hairy nature, but can easily be distinguished by their laterigrade legs, similar to those ofcrabs.Members of this family are also typically less bulky than tarantulas. They possess twoclaws,as is the case for most spiders that actively hunt their prey.If this is not enough to fully identify them, they also possess eight eyes divided into two regular rows.

Size, venom, and aggression

On average, a huntsman spider's leg-span can reach up to 15 cm (5.9 in), while their bodies measure about 1.8 cm (0.7 in) long.Like most spiders,Sparassidae use venom to immobilize prey.

There have been reports of members of various genera such asPalystes,Neosparassus,and several others inflicting severe bites on humans. The effects vary, including local swelling and pain, nausea, headache, vomiting, irregular pulse rate, and heart palpitations, indicating some systemic neurotoxin effects, especially when the bites were severe or repeated. However, the formal study of spider bites is fraught with complications, including unpredictable infections,dry bites,shock,noceboeffects, and even bite misdiagnosis by medical professionals and specimen misidentification by the general public.

It is not always clear what provokes Sparassidae to attack and bite humans and animals, but it is known that female members of this family will aggressively defend their egg-sacs and young against perceived threats.Bites from sparassids usually do not require hospital treatment.

Sound production in mating rituals

Males of the huntsman spiderHeteropoda venatoriahave recently been found to deliberately make a substrate-borne sound when they detect a chemical (pheromone) left by a nearby female of their species. The males anchor themselves firmly to the surface onto which they have crawled and then use their legs to transmit vibrations from their bodies to the surface. Most of the sound emitted is produced by strong vibrations of the abdomen. The characteristic frequency of vibration and the pattern of bursts of sound identify them to females of their species, who will approach if they are interested in mating. This sound can often be heard as a rhythmic ticking, somewhat like a quartz clock, which fades in and out and can be heard by human ears in a relatively quiet environment.

Genera

Main article:List of Sparassidae species

Isopeda villosa(lower) extricating itself from its oldexoskeleton(upper).

As of December 2022,theWorld Spider Catalogaccepted the following genera:

• AdcatomusKarsch, 1880— Venezuela, Peru
• AnaptomecusSimon, 1903— Central America, South America
• AnchonastusSimon, 1898— Cameroon, Congo
• ArandisaLawrence, 1938— Namibia
• BarylestisSimon, 1910— Africa, Asia, Europe
• BeregamaHirst, 1990— Australia, Papua New Guinea
• BerlandiaLessert, 1921— East Africa
• BhutaniellaJäger, 2000— Asia
• BorniellaGrall & Jäger, 2022— Borneo
• CaayguaraRheims, 2010— Brazil
• CarparachneLawrence, 1962— Namibia
• CebrennusSimon, 1880— Africa, Asia, Malta
• CerbalusSimon, 1897— Israel, Jordan, Egypt
• ChrosiodermaSimon, 1897— Madagascar
• ClastesWalckenaer, 1837— Indonesia, Papua New Guinea
• CuricaberisRheims, 2015— North America, Central America, Brazil
• DamastesSimon, 1880— Madagascar, Mozambique, Seychelles
• DecaphoraFranganillo, 1931— North America, Caribbean, Central America, Colombia
• DeelemanikaraJäger, 2021— Madagascar
• DefectrixPetrunkevitch, 1925— Panama
• DelenaWalckenaer, 1837— Australia, New Zealand
• DermochrosiaMello-Leitão, 1940— Brazil
• DiminutellaRheims & Alayón, 2018— Cuba
• EusparassusSimon, 1903— Asia, Africa, Europe, Peru
• ExopalystesHogg, 1914— Papua New Guinea
• ExtraordinariusRheims, 2019— Brazil
• GeminiaThorell, 1897— Myanmar
• GnathopalystesRainbow, 1899— Asia, Oceania
• GuadanaRheims, 2010— Brazil, Peru, Ecuador
• HeteropodaLatreille, 1804— Oceania, Asia, South America, Greece
• HolconiaThorell, 1877— Australia
• IrilekaHirst, 1998— Australia
• IsopedaL. Koch, 1875— Australia, Philippines, Papua New Guinea
• IsopedellaHirst, 1990— Australia, Papua New Guinea, Indonesia
• KeiliraHirst, 1989— Australia
• LeucorchestrisLawrence, 1962— Angola, Namibia
• MacrinusSimon, 1887— South America, Tobago, United States
• MartensopodaJäger, 2006— India
• MayJäger & Krehenwinkel, 2015— Namibia, South Africa
• MegaloremmiusSimon, 1903— Madagascar
• MenarikGrall & Jäger, 2022— Borneo
• MeriRheims & Jäger, 2022— South America
• MicrommataLatreille, 1804— Spain, Africa, Asia
• MicropodaGrall & Jäger, 2022— Papua New Guinea
• MicrorchestrisLawrence, 1962— Namibia
• NeosparassusHogg, 1903— Australia
• NeostasinaRheims & Alayón, 2016— Caribbean
• NolaviaKammerer, 2006— Brazil
• NungaraPinto & Rheims, 2016— Brazil, Ecuador
• OliosWalckenaer, 1837— Asia, South America, Oceania, Africa, Central America, North America, Caribbean
• OrchestrellaLawrence, 1965— Namibia
• OrigesSimon, 1897— Argentina, Peru, Ecuador
• PaenulaSimon, 1897— Ecuador
• PalystellaLawrence, 1928— Namibia
• PalystesL. Koch, 1875— Africa, India, Australia
• PanaretellaLawrence, 1937— South Africa
• PandercetesL. Koch, 1875— Asia, Oceania
• ParapalystesCroeser, 1996— South Africa
• PedianaSimon, 1880— Indonesia, Australia
• PlatnickopodaJäger, 2020— East Africa
• PleorotusSimon, 1898— Seychelles
• PolybetesSimon, 1897— South America
• PrusiasO. Pickard-Cambridge, 1892— Brazil, Mexico, Panama
• PrychiaL. Koch, 1875— Papua New Guinea, Fiji, Philippines
• PseudomicrommataJärvi, 1914— Africa
• PseudopodaJäger, 2000— Asia
• QuemediceMello-Leitão, 1942— Brazil, Argentina
• RemmiusSimon, 1897— Africa
• RhacocnemisSimon, 1897— Seychelles
• RhitymnaSimon, 1897— Asia, Africa
• SadalaSimon, 1880— South America
• SagellulaStrand, 1942— Japan, China
• SarotesiusPocock, 1898— East Africa
• SinopodaJäger, 1999— Asia
• SivalicusDyal, 1957— India
• SparianthinaBanks, 1929— South America, Tobago, Central America
• SparianthisSimon, 1880— Colombia
• SpariolenusSimon, 1880— Asia
• StaianusSimon, 1889— Madagascar
• StasinaSimon, 1877— South America, Gabon, Asia, Cuba
• StasinoidesBerland, 1922— Ethiopia
• StipaxSimon, 1898— Seychelles
• StrandiellumKolosváry, 1934— Papua New Guinea
• ThelcticopisKarsch, 1884— Asia, Oceania, Africa
• ThomasettiaHirst, 1911— Seychelles
• ThunbergaJäger, 2020— Madagascar
• TibellommaSimon, 1903— Venezuela
• TiomaniellaGrall & Jäger, 2022— Malaysia
• TychicusSimon, 1880— Philippines, Papua New Guinea, Indonesia
• TypostolaSimon, 1897— Australia, Papua New Guinea
• UaiuaraRheims, 2013— Panama, South America
• VindullusSimon, 1880— South America, Guatemala
• YiinthiDavies, 1994— Australia, Papua New Guinea
• ZachriaL. Koch, 1875— Australia

Distribution and habitat

Members of the Sparassidae are native to tropical and warm temperate regions worldwide. A few species are native to colder climates, like the green huntsman spider (Micrommata virescens) which is native toNorthernandCentral Europe.Some tropical species likeHeteropoda venatoria(Cane huntsman) andDelena cancerides(Social huntsman) have been accidentally introduced to many subtropical parts of the world, includingNew Zealand(which has no native sparassid species).

As adults, huntsman spiders do not build webs, but hunt and forage for food: their diet consists primarily of insects and otherinvertebrates,and occasionally small skinks and geckos. They live in the crevices of tree bark, but will frequently wander into homes and vehicles. They are able to travel extremely quickly, often using a springing jump while running, and walk on walls and even on ceilings. They also tend to exhibit a "cling" reflex if picked up, making them difficult to shake off and much more likely to bite. The females are fierce defenders of their egg sacs and young. They will generally make a threat display if provoked, and if the warning is ignored they may attack and bite. The egg sacs differ fairly widely among the various genera. For example, inHeteropodaspp. egg sacs are carried underneath the female's body, while in other species likePalystesandPseudomicrommataspp., females generally attach egg sacs to vegetation.

• 
  Australian sparassid egg sac hatching
• 
  Palystes castaneusegg purse

See also

• Cultural depictions of spiders (section: Modern myths and urban legends)
• List of Sparassidae species
• Spider wasp
• Table of spider families

Bibliography

• Abdullah Bayram & Sevda Özdağ (2002)."Micrommata virescens(Clerck, 1757), a new species for the spider fauna of Turkey (Araneae, Sparassidae) "(PDF).Turkish Journal of Zoology.26(3):305–307.ISSN1300-0179.(electronic).Archived(PDF)from the original on 2020-10-05.Retrieved2020-10-05.

External links

• Images and information on Australian Huntsman SpidersMinibeast Wildlife
• Information page on Huntsman Spidersfrom the Australian Museum
• Badge Huntsman Spiderat Victorian Museum
• Varieties of SparassidaeUniversity of Southern Queensland – pictures & descriptions
• Remarkable Australian Lichen HuntsmanArchived2017-03-02 at theWayback MachineatAmerican Arachnological Society
• Heteropoda venatoriaatUniversity of FloridaInstitute of Food and Agricultural Sciences
• Video of Olios sp. from Costa Rica