Paoliida
Appearance
Paoliida Temporal range:
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Fossils of Paoliida. A:Zdenekia silesiensis;B:Darekia sanguinea;C, D1,D2:Paoliidae gen. et sp. indet. | |
Scientific classification![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Cohort: | Polyneoptera |
Order: | †Paoliida Handlirsch,1906 |
Families | |
Synonyms | |
ProtopteraRasnitsyn,1977 |
Paoliidais an extinct order of winged insects that lived in the latePaleozoic.Historically, both their systematic position and composition were controversial – for instance they had been considered aspalaeodictyopterans,as basalNeoptera,or as stem-group ofPterygota– but recent studies have resolved them as thesister groupofDictyoptera.[1][2][3]
Families and genera
[edit]After Prokopet al.(2014), Guanet al.(2016) and Nel & Poschmann (2021):[1][4][5]
- †AnthracoptilidaeHandlirsch, 1922(Synonyms: StrephocladidaeMartynov, 1938;StrephoneuridaeMartynov, 1940;?AdiphlebiidaeHandlirsch, 1906)
- ?†AdiphlebiaScudder, 1885
- †AfrocladusNel, Garrouste & Prokop, 2015[6]
- †AnthracoptilusLameere, 1917
- †CarrizocladusRasnitsyn in Rasnitsynet al.,2004
- †GraticladusNovokshonov and Aristov, 2004
- †HomocladusCarpenter, 1966
- †JarmilacladusRasnitsyn and Aristov, 2004
- †LodevocladusProkopet al.,2015[7]
- †MesoptilusLameere, 1917
- †MycteroptilaRasnitsyn, 1977
- †ParacladusCarpenter, 1966
- †PruvostiaBolton, 1921
- †PseudomesoptilusGuanet al.,2016
- †RhinomaloptilaRasnitsyn, 1977
- †SpargoptilonKukalová, 1965
- †StrephocladusScudder, 1885
- †StrephoneuraMartynov, 1940
- †StrephoptilusRasnitsyn and Aristov, 2013
- †WestphaloptilusGuanet al.,2016
- †BlattinopsidaeBolton, 1925(Synonym: KlebsiellidaeHandlirsch, 1906)
- †AlienusHandlirsch, 1906
- †AvionblattinopsisQuispeet al.,2021[8]
- †BalduriellaMeunier, 1925
- †BlattinopsisGiebel, 1867
- †GlaphyrophlebiaHandlirsch, 1906
- †KlebsiellaMeunier, 1908
- †MicroblattinaScudder, 1895
- †ProtoblattiniellaMeunier, 1912
- †RhipidiopteraBrongniart, 1894
- †StephanopsisKukalová, 1958
- †WestphaloblattinopsisBéthoux and Jarzembowski, 2010
- †PaoliidaeHandlirsch, 1906(Synonyms: IdeliidaeZalessky, 1928;HerbstialidaeSchmidt, 1953;PermulidaeAristov & Storozhenko, 2011;PermotermopsidaeMartynov, 1937)
- †AcropermulaAristov and Storozhenko, 2011
- †AenigmideliaSharov, 1961
- †ArchideliaSharov, 1961
- †CarbonideliaNel, Garrouste & Jouault, 2023[9]
- †DarekiaProkopet al.,2012
- †HerbstialaSchmidt, 1952
- †HolasiciaKukalová, 1958
- †IdelinaStorozhenko, 1992
- †KemperalaBrauckmann, 1984
- †KhosarideliaStorozhenko, 1992
- †KortshakoliaSharov, 1961
- †KochopteronBrauckmann, 1984
- †MertoviaProkop and Nel, 2007
- †MicaideliaAristov 2004
- †MongoloideliaStorozhenko, 1992
- †OlinkaKukalová, 1958
- ?†PaolekiaRiek, 1976
- †PaoliaSmith, 1871
- †PaoliolaHandlirsch, 1919
- †PermotermopsisMartynov, 1937
- †PermulaHandlirsch, 1919
- †ProtoblattinaMeunier, 1909
- †PseudofouqueaHandlirsch, 1906
- †SilesiapteronProkopet al.,2013
- †SimplexpaoliaSantoset al.,2023[10]
- †SojanideliaStorozhenko, 1992
- †StenaropoditesMartynov, 1928
- †SustaiaKukalová, 1958
- †SylvideliaMartynov, 1940
- †ZdenekiaKukalová, 1958
References
[edit]- ^abJakub Prokop; Wieslaw Krzemiński; Ewa Krzemińska; Thomas Hörnschemeyer; Jan-Michael Ilger; Carsten Brauckmann; Philippe Grandcolas; André Nel (2014). "Late Palaeozoic Paoliida is the sister group of Dictyoptera (Insecta: Neoptera)".Journal of Systematic Palaeontology.12(5): 601–622.doi:10.1080/14772019.2013.823468.S2CID84407734.
- ^Legendre, Frédéric; Nel, André; Svenson, Gavin J.; Robillard, Tony; Pellens, Roseli; Grandcolas, Philippe; Escriva, Hector (22 July 2015)."Phylogeny of Dictyoptera: Dating the Origin of Cockroaches, Praying Mantises and Termites with Molecular Data and Controlled Fossil Evidence".PLOS ONE.10(7): e0130127.Bibcode:2015PLoSO..1030127L.doi:10.1371/journal.pone.0130127.PMC4511787.PMID26200914.
- ^Prokop, Jakub; Nel, André; Engel, Michael S. (2023)."Diversity, Form, and Postembryonic Development of Paleozoic Insects".Annual Review of Entomology.68(1): 401–429.doi:10.1146/annurev-ento-120220-022637.
- ^Guan, Z.; Prokop, J.; Roques, P.; Lapeyrie, J.; Nel, A. (2016)."Revision of the enigmatic family Anthracoptilidae enlightens the evolution of Palaeozoic stem-dictyopterans".Acta Palaeontologica Polonica.61(1): 71–87.doi:10.4202/app.00051.2014.
- ^André Nel; Markus J. Poschmann (2021)."Comparison of the recently described early Permian paoliid genusPermomertoviawith the 'eoblattid' genera of the families Permulidae and Permotermopsidae ".Palaeoentomology.4(1): 39–43.doi:10.11646/palaeoentomology.4.1.8.
- ^Nel, A.; Garrouste, R.; Prokop, J. (2015). "The first African Anthracoptilidae (Insecta: Paoliida) near the Permian–Triassic boundary in Kenya".Zootaxa.3925(1): 145–150.doi:10.11646/zootaxa.3925.1.10.
- ^Jakub Prokop; Jacek Szwedo; Jean Lapeyrie; Romain Garrouste; André Nel (2015). "New Middle Permian insects from Salagou Formation of the Lodève Basin in southern France (Insecta: Pterygota)".Annales de la Société Entomologique de France.New Series.51(1): 14–51.doi:10.1080/00379271.2015.1054645.S2CID87501340.
- ^Quispe, L.; Roques, P.; Garrouste, R.; Nel, A. (2021)."Carboniferous Blattinopsidae: revision ofKlebsiellaand new genus and species from Avion (Insecta, Paoliida) "(PDF).Historical Biology.34(3): 383–389.doi:10.1080/08912963.2021.1916817.S2CID234869567.
- ^Nel, A.; Garrouste, R.; Jouault, C. (2023). "New insects (Paoliida, Dictyoptera) from the Carboniferous outcrop of Tante Victoire in Var, France".Alcheringa: An Australasian Journal of Palaeontology.47(3): 305–314.doi:10.1080/03115518.2023.2258977.
- ^Santos, A. A.; Hernández-Orúe, A.; Wappler, T.; Peñalver, E.; Diez, J. B.; Nel, A. (2022). "Late Carboniferous insects from the Iberian Peninsula: State of the art and new taxa".Palaeontographica Abteilung A.326(1–6): 1–27.doi:10.1127/pala/2022/0135.S2CID251938467.