Paoliida

Paoliida Temporal range:
Fossils of Paoliida. A:Zdenekia silesiensis;B:Darekia sanguinea;C, D1,D2:Paoliidae gen. et sp. indet.
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Cohort:	Polyneoptera
Order:	†Paoliida Handlirsch,1906
Families
†Anthracoptilidae †Blattinopsidae †Paoliidae
Synonyms
ProtopteraRasnitsyn,1977

Paoliidais an extinct order of winged insects that lived in the latePaleozoic.Historically, both their systematic position and composition were controversial – for instance they had been considered aspalaeodictyopterans,as basalNeoptera,or as stem-group ofPterygota– but recent studies have resolved them as thesister groupofDictyoptera.

Families and genera

After Prokopet al.(2014), Guanet al.(2016) and Nel & Poschmann (2021):

• †AnthracoptilidaeHandlirsch, 1922(Synonyms: StrephocladidaeMartynov, 1938;StrephoneuridaeMartynov, 1940;?AdiphlebiidaeHandlirsch, 1906)
  • ?†AdiphlebiaScudder, 1885
  • †AfrocladusNel, Garrouste & Prokop, 2015
  • †AnthracoptilusLameere, 1917
  • †CarrizocladusRasnitsyn in Rasnitsynet al.,2004
  • †GraticladusNovokshonov and Aristov, 2004
  • †HomocladusCarpenter, 1966
  • †JarmilacladusRasnitsyn and Aristov, 2004
  • †LodevocladusProkopet al.,2015
  • †MesoptilusLameere, 1917
  • †MycteroptilaRasnitsyn, 1977
  • †ParacladusCarpenter, 1966
  • †PruvostiaBolton, 1921
  • †PseudomesoptilusGuanet al.,2016
  • †RhinomaloptilaRasnitsyn, 1977
  • †SpargoptilonKukalová, 1965
  • †StrephocladusScudder, 1885
  • †StrephoneuraMartynov, 1940
  • †StrephoptilusRasnitsyn and Aristov, 2013
  • †WestphaloptilusGuanet al.,2016
• †BlattinopsidaeBolton, 1925(Synonym: KlebsiellidaeHandlirsch, 1906)
  • †AlienusHandlirsch, 1906
  • †AvionblattinopsisQuispeet al.,2021
  • †BalduriellaMeunier, 1925
  • †BlattinopsisGiebel, 1867
  • †GlaphyrophlebiaHandlirsch, 1906
  • †KlebsiellaMeunier, 1908
  • †MicroblattinaScudder, 1895
  • †ProtoblattiniellaMeunier, 1912
  • †RhipidiopteraBrongniart, 1894
  • †StephanopsisKukalová, 1958
  • †WestphaloblattinopsisBéthoux and Jarzembowski, 2010
• †PaoliidaeHandlirsch, 1906(Synonyms: IdeliidaeZalessky, 1928;HerbstialidaeSchmidt, 1953;PermulidaeAristov & Storozhenko, 2011;PermotermopsidaeMartynov, 1937)
  • †AcropermulaAristov and Storozhenko, 2011
  • †AenigmideliaSharov, 1961
  • †ArchideliaSharov, 1961
  • †CarbonideliaNel, Garrouste & Jouault, 2023
  • †DarekiaProkopet al.,2012
  • †HerbstialaSchmidt, 1952
  • †HolasiciaKukalová, 1958
  • †IdelinaStorozhenko, 1992
  • †KemperalaBrauckmann, 1984
  • †KhosarideliaStorozhenko, 1992
  • †KortshakoliaSharov, 1961
  • †KochopteronBrauckmann, 1984
  • †MertoviaProkop and Nel, 2007
  • †MicaideliaAristov 2004
  • †MongoloideliaStorozhenko, 1992
  • †OlinkaKukalová, 1958
  • ?†PaolekiaRiek, 1976
  • †PaoliaSmith, 1871
  • †PaoliolaHandlirsch, 1919
  • †PermotermopsisMartynov, 1937
  • †PermulaHandlirsch, 1919
  • †ProtoblattinaMeunier, 1909
  • †PseudofouqueaHandlirsch, 1906
  • †SilesiapteronProkopet al.,2013
  • †SimplexpaoliaSantoset al.,2023
  • †SojanideliaStorozhenko, 1992
  • †StenaropoditesMartynov, 1928
  • †SustaiaKukalová, 1958
  • †SylvideliaMartynov, 1940
  • †ZdenekiaKukalová, 1958