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Taxonomic rank

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The major ranks: domain, kingdom, phylum, class, order, family, genus, and species, applied to thered fox,Vulpes vulpes.
LifeDomainKingdomPhylumClassOrderFamilyGenusSpecies
The hierarchy ofbiological classification's eight majortaxonomic ranks.Intermediate minor rankings are not shown.

Inbiology,taxonomic rankis the relative level of a group of organisms (ataxon) in an ancestral or hereditaryhierarchy.A common system of biological classification (taxonomy) consists ofspecies,genus,family,order,class,phylum,kingdom,anddomain.While older approaches to taxonomic classification were phenomenological, forming groups on the basis of similarities in appearance, organic structure and behaviour, methods based ongenetic analysishave opened the road tocladistics.

A given rank subsumes less general categories under it, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance oftraitsor features from common ancestors. The rank of anyspeciesand the description of itsgenusisbasic;which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two.[1]

Consider a particular species, thered fox,Vulpes vulpes:thespecific name or specific epithetvulpes(smallv) identifies a particular species in the genusVulpes(capitalV) which comprises all the "true" foxes. Their close relatives are all in the familyCanidae,which includes dogs, wolves, jackals, and all foxes; the next higher major rank, the orderCarnivora,includescaniforms(bears, seals, weasels, skunks, raccoons and all those mentioned above), andfeliforms(cats, civets, hyenas, mongooses). Carnivorans are one group of the hairy, warm-blooded, nursing members of the classMammalia,which are classified among animals with notochords in the phylumChordata,and with them among all animals in the kingdomAnimalia.Finally, at the highest rank all of these are grouped together with all other organisms possessingcell nucleiin the domainEukarya.

TheInternational Code of Zoological Nomenclaturedefinesrankas: "The level, for nomenclatural purposes, of ataxonin a taxonomic hierarchy (e.g. all families are for nomenclatural purposes at the same rank, which lies between superfamily and subfamily). "[2]

Main ranks[edit]

In his landmark publications, such as theSystema Naturae,Carl Linnaeusused a ranking scale limited to kingdom, class, order, genus, species, and one rank below species. Today, the nomenclature is regulated by thenomenclature codes.There are seven main taxonomic ranks: kingdom, phylum or division, class, order, family, genus, and species. In addition,domain(proposed byCarl Woese) is now widely used as a fundamental rank, although it is not mentioned in any of the nomenclature codes, and is a synonym for dominion (Latin:dominium), introduced by Moore in 1974.[3][4]

Main taxonomic ranks
Latin English
regio domain
regnum kingdom
phylum phylum(inzoology) / division (inbotany)
classis class
ordo order
familia family
genus genus
species species

A taxon is usually assigned a rank when it is given its formal name. The basic ranks are species and genus. When an organism is given a species name it is assigned to a genus, and the genus name is part of the species name.

The species name is also called abinomial,that is, a two-term name. For example, the zoological name for the human species isHomo sapiens.This is usually italicized in print or underlined when italics are not available. In this case,Homois the generic name and it is capitalized;sapiensindicates the species and it is not capitalized.

Ranks in zoology[edit]

There are definitions of the following taxonomic ranks in theInternational Code of Zoological Nomenclature:superfamily, family, subfamily, tribe, subtribe, genus, subgenus, species, subspecies.[5]

The International Code of Zoological Nomenclature divides names into "family-group names", "genus-group names" and "species-group names". The Code explicitly mentions the following ranks for these categories:[5]: §29–31 

The rules in the Code apply to the ranks of superfamily to subspecies, and only to some extent to those above the rank of superfamily. Among "genus-group names" and "species-group names" no further ranks are officially allowed. Zoologists sometimes use additional terms such asspecies group,species subgroup,species complexandsuperspeciesfor convenience as extra, but unofficial, ranks between the subgenus and species levels intaxawith many species, e.g. the genusDrosophila.(Note the potentially confusing use of "species group" as both a category of ranks as well as an unofficial rank itself.[citation needed])

At higher ranks (family and above) a lower level may be denoted by adding the prefix "infra",meaninglower,to the rank. For example,infraorder (below suborder) orinfrafamily (below subfamily).

Names of zoological taxa[edit]

  • A taxon above the rank of species has a scientific name in one part (a uninominal name).
  • A species has a name composed of two parts (a binomial name orbinomen):generic name+specific name;for exampleCanis lupus.
  • A subspecies has a name composed of three parts (a trinomial name ortrinomen): generic name + specific name +subspecific name;for exampleCanis lupus italicus.As there is only one possible rank below that of species, no connecting term to indicate rank is needed or used.

Ranks in botany[edit]

Botanical ranks categorize organisms based on their relationships. They start with Kingdom, then move to Division (or Phylum),[6]Class, Order, Family, Genus, and Species. Each rank reflects shared characteristics and evolutionary history. Understanding these ranks aids in taxonomy and studying biodiversity.

Ranks in ICN[7]: CHAPTER III 
Rank Type Suffix
kingdom (regnum) primary
subregnum further
division (divisio)
phylum (phylum)
primary ‑phyta
-mycota (fungi)
subdivisioorsubphylum further ‑phytina
-mycotina (fungi)
class (classis) primary ‑opsida (plant)
‑phyceae (algae)
-mycetes (fungi)
subclassis further ‑idae (plant)
‑phycidae (algae)
-mycetidae (fungi)
order (ordo) primary -ales
subordo further -ineae
family (familia) primary -aceae
subfamilia further ‑oideae
tribe (tribus) secondary -eae
subtribus further ‑inae
genus (genus) primary
subgenus further
section (sectio) secondary
subsectio further
series (series) secondary
subseries further
species (species) primary
subspecies further
variety (varietas) secondary
subvarietas further
form (forma) secondary
subforma further

There are definitions of the following taxonomic categories in theInternational Code of Nomenclature for Cultivated Plants:cultivar group,cultivar,grex.

The rules in the ICN apply primarily to the ranks of family and below, and only to some extent to those above the rank of family.(See alsodescriptive botanical name.)

Names of botanical taxa[edit]

Taxa at the rank of genus and above have abotanical namein one part (unitary name); those at the rank of species and above (but below genus) have a botanical name in two parts (binary name); all taxa below the rank of species have a botanical name in three parts (aninfraspecific name). To indicate the rank of the infraspecific name, a "connecting term" is needed. ThusPoa secundasubsp.juncifolia,where "subsp". is an abbreviation for "subspecies", is the name of a subspecies ofPoa secunda.[8]

Hybrids can be specified either by a "hybrid formula" that specifies the parentage, or may be given a name. For hybrids receiving ahybrid name,the same ranks apply, prefixed withnotho(Greek: 'bastard'), with nothogenus as the highest permitted rank.[9]

Outdated names for botanical ranks[edit]

If a different term for the rank was used in an old publication, but the intention is clear, botanical nomenclature specifies certain substitutions:[citation needed]

  • If names were "intended as names of orders, but published with their rank denoted by a term such as": "cohors" [Latin for "cohort";[10]see alsocohort studyfor the use of the term in ecology], "nixus", "alliance", or "Reihe" instead of "order" (Article 17.2), they are treated as names of orders.
  • "Family" is substituted for "order" (ordo) or "natural order" (ordo naturalis) under certain conditions where the modern meaning of "order" was not intended. (Article 18.2)
  • "Subfamily is substituted for" suborder "(subordo) under certain conditions where the modern meaning of" suborder "was not intended. (Article 19.2)
  • In a publication prior to 1 January 1890, if only one infraspecific rank is used, it is considered to be that of variety. (Article 37.4) This commonly applies to publications that labelled infraspecific taxa with Greek letters, α, β, γ,...

Examples[edit]

Classifications of five species follow: thefruit flyfamiliar in genetics laboratories (Drosophila melanogaster),humans(Homo sapiens), thepeasused byGregor Mendelin his discovery ofgenetics(Pisum sativum), the "fly agaric" mushroomAmanita muscaria,and the bacteriumEscherichia coli.The eight major ranks are given in bold; a selection of minor ranks are given as well.

Rank Fruit fly Human Pea Fly agaric E. coli
Domain Eukarya Eukarya Eukarya Eukarya Bacteria
Kingdom Animalia Animalia Plantae Fungi Pseudomonadati[11]
Phylumordivision Arthropoda Chordata Magnoliophyta(Tracheophyta) Basidiomycota Pseudomonadota
Subphylumor subdivision Hexapoda Vertebrata Magnoliophytina (Euphyllophytina) Agaricomycotina
Class Insecta Mammalia Magnoliopsida(Equisetopsida) Agaricomycetes Gammaproteobacteria
Subclass Pterygota Theria Rosidae(Magnoliidae) Agaricomycetidae
Superorder Panorpida Euarchontoglires Rosanae
Order Diptera Primates Fabales Agaricales Enterobacterales
Suborder Brachycera Haplorrhini Fabineae Agaricineae
Family Drosophilidae Hominidae Fabaceae Amanitaceae Enterobacteriaceae
Subfamily Drosophilinae Homininae Faboideae Amanitoideae
Tribe Drosophilini Hominini Fabeae
Genus Drosophila Homo Pisum Amanita Escherichia
Species D. melanogaster H. sapiens P. sativum A. muscaria E. coli
Table notes
  • In order to keep the table compact and avoid disputed technicalities, some common and uncommon intermediate ranks are omitted. For example, themammalsof Europe, Africa, and upper North America[a]are in classMammalia,legionCladotheria,sublegionZatheria,infralegionTribosphenida,subclassTheria,cladeEutheria,cladePlacentalia– but only Mammalia and Theria are in the table. Legitimate arguments might arise if the commonly used clades Eutheria and Placentalia were both included, over which is the rank "infraclass" and what the other's rank should be, or whether the two names are synonyms.
  • The ranks of higher taxa, especially intermediate ranks, are prone to revision as new information about relationships is discovered. For example, theflowering plantshave been downgraded from a division (Magnoliophyta) to a subclass (Magnoliidae), and the superorder has become the rank that distinguishes the major groups of flowering plants.[12]The traditional classification of primates (class Mammalia, subclass Theria, infraclass Eutheria, order Primates) has been modified by new classifications such as McKenna and Bell (class Mammalia, subclass Theriformes, infraclass Holotheria) with Theria and Eutheria assigned lower ranks between infraclass and the order Primates.(Seemammal classificationfor details.)These differences arise because there are few available ranks and many branching points in the fossil record.
  • Within species further units may be recognised. Animals may be classified into subspecies (for example,Homo sapiens sapiens,modern humans) ormorphs(for exampleCorvus corax variusmorphaleucophaeus,the pied raven). Plants may be classified into subspecies (for example,Pisum sativumsubsp.sativum,the garden pea) or varieties (for example,Pisum sativumvar.macrocarpon,snow pea), with cultivated plants getting acultivarname (for example,Pisum sativumvar.macrocarpon'Snowbird'). Bacteria may be classified bystrains(for exampleEscherichia coliO157:H7,a strain that can causefood poisoning).

Terminations of names[edit]

Taxaabove the genus level are often given names based on thetype genus,with a standard termination. The terminations used in forming these names depend on the kingdom (and sometimes the phylum and class) as set out in the table below.

Pronunciations given are themost Anglicized.More Latinate pronunciations are also common, particularly/ɑː/rather than//for stresseda.

Rank BacteriaandArchaea[13] Embryophytes(Plants) Algae(Diaphoretickes) Fungi Animals Viruses[14]
Realm -viria
Subrealm -vira
Kingdom -ati[15] -virae
Subkingdom -viretes
Division/phylum -ota[16] -ophyta[17]/ˈɒfətə,ə(ˈ)ftə/ -mycota/mˈktə/ -viricota/vɪrəˈktə/
Subdivision/subphylum -phytina[17]/fəˈtnə/ -mycotina/mkəˈtnə/ -viricotina/vɪrəkəˈtnə/
Class -ia/iə/ -opsida/ˈɒpsədə/ -phyceae/ˈfʃ/ -mycetes/mˈstz/ -viricetes/vɪrəˈstz/
Subclass -idae/əd/ -phycidae/ˈfɪsəd/ -mycetidae/mˈsɛtəd/ -viricetidae/vɪrəˈsɛtəd/
Superorder -anae/ˈn/
Order -ales/ˈlz/ -ida/ədə/or -iformes/ə(ˈ)fɔːrmz/ -virales/vˈrlz/
Suborder -ineae/ˈɪn/ -virineae/vəˈrɪn/
Infraorder -aria/ˈɛəriə/
Superfamily -acea/ˈʃə/ -oidea/ˈɔɪdə/
Epifamily -oidae/ˈɔɪd/
Family -aceae/ˈʃ/ -idae/əd/ -viridae/ˈvɪrəd/
Subfamily -oideae/ˈɔɪd/ -inae/ˈn/ -virineae/vɪˈrɪn/
Infrafamily -odd/ɒd/[18]
Tribe -eae// -ini/ˈn/
Subtribe -inae/ˈn/ -ina/ˈnə/
Infratribe -ad/æd/or -iti/ˈti/
Genus -virus
Subgenus -virus
Table notes
  • In botany and mycology names at the rank of family and below are based on the name of a genus, sometimes called thetype genusof that taxon, with a standard ending. For example, the rose family,Rosaceae,is named after the genusRosa,with the standard ending "-aceae" for a family. Names above the rank of family are also formed from a generic name, or are descriptive (likeGymnospermaeorFungi).
  • For animals, there are standard suffixes for taxa only up to the rank of superfamily.[19]Uniform suffix has been suggested (but not recommended) inAAAS[20]as-ida/ɪdə/for orders, for example; protozoologists seem to adopt this system. Many metazoan (higher animals) orders also have such suffix, e.g.HyolithidaandNectaspida (Naraoiida).
  • Forming a name based on a generic name may be not straightforward. For example, thehomohas the genitivehominis,thus the genusHomo(human) is in theHominidae,not "Homidae".
  • The ranks of epifamily, infrafamily and infratribe (in animals) are used where the complexities of phyletic branching require finer-than-usual distinctions. Although they fall below the rank of superfamily, they are not regulated under the International Code of Zoological Nomenclature and hence do not have formal standard endings. The suffixes listed here are regular, but informal.[21]
  • In virology, the formal endings for taxa ofviroids,ofsatellite nucleic acids,and ofviriformsare similar to viruses, only-vir-is replaced by-viroid-,-satellit-and-viriform-.[14]The extra levels of realm and subrealm end with-viriaand-virarespectively.[14]

All ranks[edit]

There is an indeterminate number of ranks, as a taxonomist may invent a new rank at will, at any time, if they feel this is necessary. In doing so, there are some restrictions, which will vary with thenomenclature codethat applies.[citation needed]

The following is an artificial synthesis, solely for purposes of demonstration of relative rank (but see notes), from most general to most specific:[22]

Significance and problems[edit]

Ranks are assigned based on subjective dissimilarity, and do not fully reflect the gradational nature of variation within nature. In most cases, higher taxonomic groupings arise further back in time: not because the rate of diversification was higher in the past, but because each subsequent diversification event results in an increase of diversity and thus increases the taxonomic rank assigned by present-day taxonomists.[27]Furthermore, some groups have many described species not because they are more diverse than other species, but because they are more easily sampled and studied than other groups.[citation needed]

Of these many ranks, the most basic is species. However, this is not to say that a taxon at any other rank may not be sharply defined, or that any species is guaranteed to be sharply defined. It varies from case to case. Ideally, a taxon is intended to represent aclade,that is, thephylogenyof the organisms under discussion, but this is not a requirement.[citation needed]

A classification in which all taxa have formal ranks cannot adequately reflect knowledge about phylogeny. Since taxon names are dependent on ranks in traditional Linnaean systems of classification, taxa without ranks cannot be given names. Alternative approaches, such as usingcircumscriptional names,avoid this problem.[28][29]The theoretical difficulty with superimposing taxonomic ranks over evolutionary trees is manifested as theboundary paradoxwhich may be illustrated by Darwinian evolutionary models.

There are no rules for how many species should make a genus, a family, or any other higher taxon (that is, a taxon in a category above the species level).[30][31]It should be a natural group (that is, non-artificial, non-polyphyletic), as judged by a biologist, using all the information available to them. Equally ranked higher taxa in different phyla are not necessarily equivalent (e.g., it is incorrect to assume that families of insects are in some way evolutionarily comparable to families of mollusks).[31]For animals, at least thephylumrank is usually associated with a certainbody plan,which is also, however, an arbitrary criterion.[citation needed]

Mnemonic[edit]

There are several acronyms intended to help memorise the taxonomic hierarchy, such as "King Phillip came over for great spaghetti".[32](Seetaxonomy mnemonic.)

See also[edit]

Footnotes[edit]

  1. ^TheVirginia opossumis an exception.

References[edit]

  1. ^"International Code of Nomenclature for algae, fungi, and plants – Melbourne Code".IAPT-Taxon.org.2012. Articles 2 and 3.Archivedfrom the original on 10 June 2019.Retrieved28 April2013.
  2. ^International Commission on Zoological Nomenclature (1999),International Code of Zoological Nomenclature. Fourth Edition,International Trust for Zoological Nomenclature,archivedfrom the original on 21 May 2019,retrieved12 April2015
  3. ^Moore, R. T. (1974)."Proposal for the recognition of super ranks"(PDF).Taxon.23(4): 650–652.doi:10.2307/1218807.JSTOR1218807.Archived(PDF)from the original on 6 October 2016.Retrieved5 October2016.
  4. ^Luketa, S. (2012)."New views on the megaclassification of life"(PDF).Protistology.7(4): 218–237. Archived fromthe original(PDF)on 2 April 2015.
  5. ^abInternational Commission on Zoological Nomenclature (1999). "Glossary".International Code of Zoological Nomenclature.London: The International Trust for Zoological Nomenclature.ISBN0-85301-006-4.Archivedfrom the original on 3 September 2021.Retrieved10 November2023.
  6. ^jibran, jibran."Log In ‹ Information metBotanical Ranks: Understanding Taxonomic Classification" Meta Description: Explore the hierarchical structure of botanical classification, from Kingdom to Species, essential for understanding plant diversity and evolution. a description — WordPress ".dev-information-meta-descrition.pantheonsite.io.Retrieved7 May2024.
  7. ^"International Code of Nomenclature for algae, fungi, and plants – Melbourne Code".IAPT-Taxon.org.2012.Archivedfrom the original on 10 October 2020.Retrieved28 April2013.
  8. ^"International Code of Nomenclature for algae, fungi, and plants – Melbourne Code".IAPT-Taxon.org.2012. Articles 4.2 and 24.1.Archivedfrom the original on 3 August 2018.Retrieved3 August2018.
  9. ^"International Code of Nomenclature for algae, fungi, and plants – Melbourne Code".IAPT-Taxon.org.2012. Article 3.2, and Appendix 1, Articles H.1–3.Archivedfrom the original on 10 October 2020.Retrieved28 April2013.
  10. ^Stearn, W.T. 1992.Botanical Latin: History, grammar, syntax, terminology and vocabulary, Fourth edition.David and Charles.
  11. ^Göker, Markus; Oren, Aharon (22 January 2024)."Valid publication of names of two domains and seven kingdoms of prokaryotes".International Journal of Systematic and Evolutionary Microbiology.74(1). Society for General Microbiology.doi:10.1099/ijsem.0.006242.ISSN1466-5034.PMID38252124.Retrieved7 June2024.
  12. ^Chase, M. W.; Reveal, J. L. (2009), "A phylogenetic classification of the land plants to accompany APG III",Botanical Journal of the Linnean Society,161(2): 122–127,doi:10.1111/j.1095-8339.2009.01002.x
  13. ^Oren, Aharon(2023)."International Code of Nomenclature of Prokaryotes. Prokaryotic Code (2022 Revision)".International Journal of Systematic and Evolutionary Microbiology.73(5a).doi:10.1099/ijsem.0.005585.hdl:10261/338243.PMID37219928.
  14. ^abcde"ICTV Code. Section 3.IV, § 3.23; section 3.V, §§ 3.27-3.28."International Committee on Taxonomy of Viruses.October 2018.Archivedfrom the original on 6 July 2022.Retrieved28 November2018.
  15. ^Oren, Aharon (1 November 2023). "Emendation of Principle 8, Rules 5b, 8, 15, 33a, and Appendix 7 of the International Code of Nomenclature of Prokaryotes to include the categories of kingdom and domain".International Journal of Systematic and Evolutionary Microbiology.73(11).doi:10.1099/ijsem.0.006123.PMID37909283.
  16. ^Whitman, William B.; Oren, Aharon; Chuvochina, Maria; da Costa, Milton S.; Garrity, George M.; Rainey, Fred A.; Rossello-Mora, Ramon; Schink, Bernhard; Sutcliffe, Iain; Trujillo, Martha E.; Ventura, StefanoYR 2018 (2018)."Proposal of the suffix –ota to denote phyla. Addendum to 'Proposal to include the rank of phylum in the International Code of Nomenclature of Prokaryotes'".International Journal of Systematic and Evolutionary Microbiology.68(3): 967–969.doi:10.1099/ijsem.0.002593.ISSN1466-5034.PMID29458499.{{cite journal}}:CS1 maint: numeric names: authors list (link)
  17. ^ab"International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code)".IAPT-Taxon.org.2018. Article 16.Archivedfrom the original on 19 December 2018.Retrieved19 December2018.
  18. ^For example, thechelonianinfrafamilies Chelodd (Gaffney & Meylan 1988: 169) and Baenodd (ibid.,176).
  19. ^ICZN article 29.2
  20. ^Pearse, A.S. (1936)Zoological names. A list of phyla, classes, and orders, prepared for section F, American Association for the Advancement of Science.Archived15 November 2021 at theWayback MachineAmerican Association for the Advancement of Science, p. 4
  21. ^As supplied by Gaffney & Meylan (1988).
  22. ^For the general usage and coordination of zoological ranks between the phylum and family levels, including many intercalary ranks, see Carroll (1988). For additional intercalary ranks in zoology, see especially Gaffney & Meylan (1988); McKenna & Bell (1997); Milner (1988); Novacek (1986, cit. in Carroll 1988: 499, 629); andPaul Sereno's 1986 classification ofornithischiandinosaurs as reported in Lambert (1990: 149, 159). For botanical ranks, including many intercalary ranks, see Willis & McElwain (2002).
  23. ^abcdThese are movable ranks, most often inserted between the class and the legion or cohort. Nevertheless, their positioning in the zoological hierarchy may be subject to wide variation. For examples, see theBenton classification of vertebratesArchived16 January 2019 at theWayback Machine(2005).
  24. ^abcdIn zoological classification, the cohort and its associated group of ranks are inserted between the class group and the ordinal group. The cohort has also been used between infraorder and family insaurischiandinosaurs (BentonArchived16 January 2019 at theWayback Machine2005). In botanical classification, the cohort group has sometimes been inserted between the division (phylum) group and the class group: see Willis & McElwain (2002: 100–101), or has sometimes been used at the rank of order, and is now considered to be an obsolete name for order: See International Code of Nomenclature for algae, fungi, and plants, Melbourne Code 2012, Article 17.2.
  25. ^abcdeThe supra-ordinal sequence gigaorder–megaorder–capaxorder–hyperorder (and the microorder, in roughly the position most often assigned to the parvorder) has been employed in turtles at least (Gaffney & Meylan 1988), while the parallel sequence magnorder–grandorder–mirorder figures in recently influential classifications of mammals. It is unclear from the sources how these two sequences are to be coordinated (or interwoven) within a unitary zoological hierarchy of ranks. Previously, Novacek (1986) and McKenna-Bell (1997) had inserted mirorders and grandorders between the order and superorder, but Benton (2005) now positions both of these ranks above the superorder.
  26. ^Additionally, the termsbiovar,morphovar,phagovar,andserovardesignate bacterialstrains(genetic variants) that are physiologically or biochemically distinctive. These are not taxonomic ranks, but are groupings of various sorts which may define a bacterial subspecies.
  27. ^Gingerich, P. D. (1987). "Evolution and the fossil record: Patterns, rates, and processes".Canadian Journal of Zoology.65(5): 1053–1060.doi:10.1139/z87-169.
  28. ^Kluge, N.J. (1999). "A system of alternative nomenclatures of supra-species taxa. Linnaean and post-Linnaean principles of systematics".Entomological Review.79(2): 133–147.
  29. ^Kluge, N.J. (2010)."Circumscriptional names of higher taxa in Hexapoda".Bionomina.1(1): 15–55.doi:10.11646/bionomina.1.1.3.
  30. ^Stuessy, T.F. (2009).Plant Taxonomy: The Systematic Evaluation of Comparative Data.2nd ed. Columbia University Press, p. 175.
  31. ^abBrusca, R.C. & Brusca, G.J. (2003).Invertebrates.2nd ed. Sunderland, Massachusetts: Sinauer Associates, pp. 26–27.
  32. ^Evans, Rod L. (2007).Every Good Boy Deserves Fudge: The Book of Mnemonic Devices.Penguin.ISBN978-1-4406-2207-6.Archivedfrom the original on 23 November 2023.Retrieved23 November2023.

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