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Title Director Format Info
Looney Tunes
Mad as a Mars Hare		 Chuck Jones Cartoon Series Bugs Bunnyis turned into a "Neanderthal Rabbit" after getting hit by a ray from a time-projector gun byMarvin the Martian.
Quest for Fire Jean-Jacques Annaud 1981 film features Neanderthals and a Cro-Magnon attempting to carry a vessel containing fire to the Neanderthal's tribe.
Clan of the Cave Bear Michael Chapman 1986 film novel by Jean M. Auel about prehistoric times. It is the first book in the Earth's Children book series
Iceman Fred Schepisi 1984 film from a screenplay written by John Drimmer, depicts a frozen Neanderthal coming to life again in the 1980s at an arctic research station.
Ghost Light Andrew Cartmel 1989 film an episode in the television seriesDoctor Who,a Neanderthal called "Nimrod" is thebutlerof aVictorian erahousehold.
Night at the Museum Shawn Levy 2006 film four Neanderthals were put on display in theAmerican Museum of Natural History.Anancient Egyptiantablet, the Tablet of Akhemrah, causes everything on display in the museum to come to life at night. The Neanderthals showed an interest in fire after it was shown to them by the night guard, Larry Daley.
The Real Adventures of Jonny Quest Sherry Gunther TV series the mythicalyetisare stated to be arelictpopulation of Neanderthals.
Dinosaurs Brian Henson TV series Generic "cavemen" have appeared in multiple episodes notablyseason 3episodes, "The Discovery," and "Charlene and Her Amazing Humans."
You Can't Do That on Television Geoffrey Darby TV program A Neanderthal-like family was a frequent recurring sketch in the children's show, In keeping with the theme of that particular episode, the sketch often parodied modern issues with coarse, overbearing parents outside of a pre-historic cave setting.[1]
GEICO Cavemen Joe Lawson advertisement trademarkedcharacters in a series of television advertisements for theauto insurancecompanyGEICOthat have aired from 2004 to present, featuring Neanderthal-like cavemen in a modern setting
The Croods Chris Sanders
Kirk DeMicco		 2013 animated film features the titular family as they embark on a journey to find a new home along with aCro-Magnonboy who has mastered fire and other "technologies" they had never previously encountered.
Walking with Beasts Tim Haines Documentary One is charged by awoolly rhinoceros,but escapes, in part because of his stocky constitution. The climax of the episode is when the clan of Neanderthals attack the herd ofmammothas they turn back to the north.
Ao: The Last Hunter
Ao, le dernier Néandertal		 Jacques Malaterre prehistoric film Ao is the protagonist in a 2010 French prehistoric film[2]

The contemporary perception of Neanderthals and their stereotypical portrayal has its origins in 19th century Europe. Naturalists and anthropologists were confronted with an increasing number of fossilized bones that that did not match any knowntaxon.Carl Linnaeuss1758Systema Naturaein which he hadHomo sapiensintroduced as a species without diagnosis and description, was the authoritative encyclopedia of the time. The notion of extinct species was unheard of and if so, would have contradicted the paradigm of the immutability of species and the physical world, which was the infallible product of a single and deliberate act of a creator god. Most scholars simply declared the early Neanderthal fossils to be representatives of early "races" of modern man.Thomas Henry Huxley,a future supporter ofDarwin'stheory of evolution,saw in theEngis 2fossil a "man of low degree of civilization". The discovery in theNeandertalhe interpreted as to be within the range of variation of modern humans.[3]

In mid 19th century Germany biological sciences were dominated byRudolf Virchow,who described the bones as a "remarkable individual phenomenon" and as "plausible individual deformation".[4]This statement is the reason why the characteristics of the Neanderthals were perceived as a form of pathological skeleton change of modern man in German-speaking countries for many years to come.

August Franz Josef Karl Mayer, an associate of Virchow emphasizes disease, prolonged pain and struggle on comparison with modern human features.[5]"He confirmed the Neanderthal's rachitic changes in bone development[...]. Mayer argued among other things, that the thigh - and pelvic bones of Neanderthal man were shaped like those of someone who had spent all his life on horseback. The broken right arm of the individual had only healed very badly and the resulting permanent worry lines about the pain were the reason for the distinguished brow ridges. The skeleton was, he speculated, that of a mounted RussianCossack,who had roamed the region in 1813/14 during the turmoils of thewars of liberationfromNapoleon."[4]


Arthur Keithof Britain andMarcellin Bouleof France, were both senior members of their respective national paleontological institutes and among the most eminent paleoanthropologists of the early 20th century. Both men argued that this "primitive" Neanderthal could not be a direct ancestor of modern man. As a result the museum's copy of the almost complete Neanderthal fossil ofLa Chapelle-aux-Saintswas inaccurately mounted in an exaggerated crooked pose with a deformed and heavily curved spine and legs buckled.[6][7][8][9]


Northern Canada
Canadian Arctic Tundra
Northern MainlandCanadian Arctic Archipelago
Southern ArcticNorthern ArcticArctic Cordillera
Baffin and Torngat MountainsEllesmere and Devon Islands Ice Caps
Gulf of Boothia and Foxe Basin PlainsBanks Island and Amundsen Gulf LowlandsVictoria Island LowlandsBaffin UplandsFoxe UplandsLancaster and Borden Peninsula PlateausParry Islands PlateauEllesmere Mountains and Eureka HillsSverdrup Islands Lowland
Amundsen PlainsAberdeen PlainsCentral Ungava Peninsula and Ottawa and Belcher IslandsQueen Maud Gulf and Chantrey Inlet LowlandsMackenzie and Selwyn MountainsPeel River and Nahanni Plateaus

Source:[10]


Neanderthal
Temporal range:Middle–Late Pleistocene
Neanderthal skull atLa Chapelle-aux-Saints
Scientific classificationEdit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Suborder: Haplorhini
Infraorder: Simiiformes
Family: Hominidae
Subfamily: Homininae
Tribe: Hominini
Genus: Homo
Species:
H. neanderthalensis
Binomial name
Homo neanderthalensis
King,1864
Range ofHomo neanderthalensis,that may extend to include theOkladnikovCave/Altai Mountainsand Mamotnaia/Ural Mountains
Synonyms

Homo mousteriensis hauseri[11]
Homo sapiens neanderthalensis[12]
Palaeoanthropus neanderthalensis[13]

NeanderthalsorNeandertalsUK:/niˈændərˌtɑːl/,usalso/neɪ/-, -/ˈɑːndər/-, -/ˌtɔːl/,-/ˌθɔːl/)[14][15]are an extinctspeciesor subspecies of thegenusHomo,that was a close relative ofmodern humans.Both species occupied a common habitat inEuropefor several thousand years. Research has so far no universally accepted conclusive explanation as to what caused the Neanderthal's extinction between 40.000 and 28.000 years ago.[16]

Neanderthal classification has been subject to debate since the discovery of the fdo to this day.[17]Currently two formal species names reflect the ongoing controversy -Homo neanderthalensisthe binominal name of a distinctspeciesandHomo sapiens neanderthalensis,that classifies Neanderthals as asubspeciesofHomo sapiens.[18][19][20]

Sharing a common ancestor of thegenusHomowithHomo sapiens,who evolved independently in Africa,[21]the specie's specific morphology developed inEurasiaduring the last 350.000 years, reaching its final, "classical Neanderthal" form approximately 130.000 years ago.[22]

Neanderthals left a rich fossil record in thelimestonecaves ofEurasia,fromWestern Europeto theAltai MountainsinCentral Asiaand theUral Mountainsin the North to theLevantin the South.[23]Discoveries include culturalassemblagesand Neanderthals are associated with theMousterian culture,which first appeared approximately 160.000 years ago.[24][25][26]

Generally small and widely dispersed fossil sites suggest, that Neanderthals lived in less numerous and socially more isolated groups than contemporaryHomo sapiens.Tools such as Mousterian flint stone flakes andLevallois pointsare remarkably sophisticated from the outset, yet they have a slow rate of variability and general technological inertia is noticeable during the entire fossil period. Artefacts are of utilitarian nature, symbolic behavioral traits are undocumented before the arrival of modern humans in Europe around 40.000 to 35.000 years ago.[27][28]

The Neanderthal genome sequence was successfully generated in 2010 as the central assignment for theNeanderthal genome project.[29]The DNA structure proved to be 99.7% identical with anatomically modern humans.[30]Subsequently the genetic relationship is being investigated[31][32]and ongoing research has yet helped to interpret and verify kinship lines and interbreeding theories with modern humans and other species of the genusHomoand resulting evolutionary implications.[33][34][35][36]

In 2013 researchers announced that the assemblages of at least 40 widely scattered archaeological sites suggest that Neanderthals intentionally buried their dead.[37]In 2016 a team of researchers found ring structures made ofstalagmitesfragments inside theBruniquel Cave,south-western France around 176.000 years old, which were attributed to early Neanderthals. These discoveries might revive the debate on the degree of advancement of Neanderthal's social structure.[38]

Etymology[edit]

The species was named by geologistWilliam Kingin his 1864 species descriptionThe Reputed Fossil Man of the Neanderthalafter having introduced the termHomo Neanderthalensis Kingone year earlier giving a lecture at theBritish Association for the Advancement of Science.[39][40][41]The type specimenNeanderthal 1was only discovered in 1856 in alimestonegrotto in the Neandertal (Neander-valley, spelled Neanderthal before the 1901 German spelling reform), 12 km (7.5 mi) east ofDüsseldorf,Germany.[42]

Discovery[edit]

Neanderthal excavation site andNeander ValleyinNorth Rhine-Westphalia,Germany

The first recorded Neanderthal fossil of a partial skull and some associated fragments, later dubbedEngis 2was discovered in 1829 by Dutch physician and naturalistPhilippe-Charles Schmerlingin theAwirsCave, just north of theBelgianmunicipalityEngis[43]and professionally described in 1833.[44][45]The second fossilGibraltar 1,was discovered by Captain Edmund Flint in theForbes' Quarryof Gibraltar in 1848. Both findings received only little attention and were misunderstood as to represent early forms or pathologically developed, abnormal human individuals.[46]A universal definition of species-specific characteristics of species in the genus Homo did not exist in the early 19th century. Methodical recognition of distinct and extinct species only began afterCharles Darwin's publication ofOn the Origin of Speciesin 1859.[47]

The 1856 in alimestonequarry - theKleine Feldhofer Grottein the Neandertal discoveredtype specimenconsisted of a skull cap, two femora, three bones from the right arm, two from the left arm, part of the left ilium, fragments of a scapula, and ribs. These quickly came into possession of local teacher and amateur naturalistJohann Carl Fuhlrott,who studied the fossil in co-operation with anatomistHermann Schaaffhausen.The jointly made publication in 1857 represents the beginning ofpaleoanthropologyas a scientific discipline.[48][49]The scholarly community remained divided for another 30 years until the discovery of two well preserved and almost complete Neanderthal skeletons in Spy/Belgium in 1886 lead to general recognition of the species.[50]

Origin[edit]

Altamura Man (between 128.000 and 187.000 years old) is the oldest fossil of the Neanderthal lineage, that has yielded a reliable DNA sequence.[51]Older specimen are until recently interpreted and classified based on morphology and metric analyses. 430.000 years old skull fossils recovered from theSima de los Huesos cavehave proto-Neanderthal features, such as "the beginning of a prominent brow ridge, a distinctive jaw shape and patterns of cusps on the teeth, [but] lack other traits that define the species — notably a large cranium".[52][53][54]DNA tests confirm that these skulls represent early Neanderthals.[55]Anthropologists debate over whether the species evolved fromHomo heidelbergensis,or from what some researchers call "a controversial species" -Homo antecessor.[56]

"Not quite Neanderthals, the Sima de los Huesos humans probably represent one of many small, isolated groups that dotted the European continent at the time, notes Jean-Jacques Hublin, a palaeoanthropologistat at the Max Planck Institute for Evolutionary Anthropology in Leipzig, Germany. Such isolation is ideal for anatomical features to be established by chance, and this genetic drift could explain many of the attributes that would later define Neanderthals."

Fossils found inTourville-la-Rivière,[57]the 400.000 year oldTautavel Man,the 600.000 year old mandibleMauer 1and thePetralona skullhave all been attributed to the Neanderthal lineage, although only based on morphological and metric analyses.[58]

Classification[edit]

Scientists to this day debate over whether Neanderthals should be classified as a distinct species -Homo neanderthalensis- or asHomo sapiens neanderthalensis,the latter placing Neanderthals as a subspecies ofH. sapiens.[59][60]During the early 20th century the prevailing view has been heavily influenced byArthur KeithandMarcellin Boule,who wrote the first scientific description of a nearly complete Neanderthal skeleton.[61]Senior members of their respective national paleontological institutes and among the most eminent paleoanthropologists of their time,[62][63]both men argued that this "primitive" Neanderthal could not be a direct ancestor of modern man. This idea was reflected in an erroneous and inaccurate reconstruction of the Neanderthal findings ofLa Chapelle-aux-Saints,mounted in a crooked pose with a deformed and heavily curved spine and legs buckled.[64]

During the 1930s scholarsErnst Mayr,George Gaylord SimpsonandTheodosius Dobzhanskyreinterpreted the existing fossil record and came to different conclusions.[65]Neanderthal man was classified asHomo sapiens neanderthalensis- an early subspecies of what was now calledHomo sapiens sapiens.The obviously unbroken succession of fossil sites of both subspecies in Europe was considered evidence that there was a slow and gradual evolutionary transition from Neanderthals to modern humans. Contextual interpretations of similar excavation sites in Asia lead to thehypothesis of multiregional origin of modern manin the 1980s.[66]

Contemporary views[edit]

Current scientific ideas hold that both species evolved from a common African ancestor -Homo erectus.Fossil and tool finds support the idea that during the first migration wave of around 2 million years ago small groups ofHomo erectusleft Africa via theLevant,theBlack Sea areaintoGeorgia(fossils of Dmanisi) and possibly via North-western Africa towards southern Spain. The 1.2 million years old fragmented finds in Spain are referred to asHomo antecessorby their discoverers and considered to be the ancestors of Neanderthal. This interpretation, however, is controversial.

Some 600,000 years ago a second propagation wave of AfricanHomo erectustook place. Skulls from that period found in Spain for example, suggest a brain volume of between 1100 ccm to 1450 ccm. The brain volume of fossils from the first propagation wave is however estimated only to be slightly above 1000 ccm.Homo erectusthat arrived in Europe in the second wave of colonization developed into Neanderthals through the intermediateHomo heidelbergensis.Since about 200,000 years ago in Africa the so-calledearly anatomically modern humansdescended from the localHomo erectusthrough the intermediateHomo rhodesiensis.Evidence frommitochondrial DNAstudies has been interpreted as evidenceNeanderthalswere not a subspecies ofH. sapiens.[67]Others, for exampleUniversity of CambridgeProfessorPaul Mellars,say "no evidence has been found of cultural interaction".[68]

Debated remains the period when the Neanderthal lineage separated from the lineage that lead to modern man. In 2010 a period between 440,000 - 270,000 years B.P. was calculated by themolecular clockmethod. However, the reliability of molecular clocks is questionable as dating determined bystratigraphicmethods varies considerably from those determined using the molecular clock. In 2012 recalculation of the mutation rates revealed evidence of a much earlier separation between 800,000 and 400,000 years ago.


Timeline of discovery and research[edit]

Neanderthals are afterH. sapiensthe best documented hominin species as they habitually occupied limestone caves and rock shelters, locations with a high fossilization and preservation ratio for bones, artefacts and fire places. The bones of over 400 specimen have been found since the early 19th century.[69]


Timeline and locations of discovery, specimen age, associated research and data
Year Discovery Research Fossil age Location Notability Image
1829 Engis 2byPhilippe-Charles Schmerling described in 1833 but not specified Awirs Cave,Belgium first fossil discovery
1848 Gibraltar 1Skull by Captain Edmund Flint Forbes' QuarryGibraltar
1856 Neanderthal 1byJohann Karl Fuhlrott described as a separate species and published in 1857, but not universally recognized 40.000 BP. Neandertal, Germany first paleo-anthropologic publication,
1864 William Kingintroduces the species nameHomo neanderthalensis
1880

The Šipka cave mandibleby Karel Jaroslav Maška

first discovery of Neanderthal remains in their cultural context Šipka,Czech Republic firstMousteriantools discovered
1886

The Spy Skeletonsby Marcel de Puydt and Max Lohest

discovery lead to international scientific recognition of the existence of more than one human species Spy Cave,Belgium cave site was also occupied byH. sapiens,who might have lived alongside Neanderthal
1899

The Krapina FossilsbyDragutin Gorjanović-Kramberger

over eight hundred fossil remains discovered, first evidence for morphological variations Hušnjakovo Hill,Krapina,Croatia fossils prove Neanderthal evolution
1908

La Chapelle-aux-Saints 1by Jean and Amédée Bouyssonie, L. Bardon[70]

discovery of over 1,000 pieces of stone (flint) and bone tools, Mousterian assemblages 60,000 years BP. La Chapelle-aux-Saints,France first recognized burial of a Neanderthal
1909

La FerrassieSkull by Denis Peyrony and Louis Capitan

Eight buried Neanderthals have been found in La Ferrassie, including infants and two fetuses in a cave flanked by two rock shelters 70–50,000 years BP. Savignac-de-Miremont,Dordogne,France. The La Ferrassie 1 skull is the largest and most complete Neanderthal skull ever found
1909

Le Moustierskull with a large nasal cavity and a somewhat less developed brow ridge and occipital bun, as might be expected in a juvenile.

The site are two rock shelters near a village in the Dordogne, first excavated from 1863 by the EnglishmanHenry Christyand the FrenchmanÉdouard Lartet. 45,000 years BP. Peyzac-le-Moustier,Dordogne,France. TheMousteriantool culture is named after this site
1926 Skull fragments ofGibraltar 2,a four-year-old Neanderthal girl, discovered byDorothy Garrod Devil's Tower cave,Gibraltar
1953–1957 Ralph Soleckiuncovered nine Neanderthal skeletons in Shanidar Cave, Kurdistan A Mousterian layer which contained hundreds of stone tools including points, side-scrapers, flakes and bones, good condition of fossils yields data and information on individual specimen. 35,000 to 65,000 years BP. Shanidar,Irak Shanidar 1, survived severe injuries attributed to care from other members of his band, Shanidar 4, lay beside a flower that might be explained as evidence of burial rituals - or animal action.
1961-1964 Amud 1- the skeleton of an adult male about 25 years old by University of Tokyo expeditions Amud 1 has a cranial capacity of about 1,740 cubic cm (106 cubic inches), which is significantly larger than the average capacity of modern humans. Nonetheless, the skull has Neanderthal features, with browridges and a receding forehead. 50,000 to 70,000 years BP. Nahal AmudUpper Galilee evidence of Neanderthal habitation and Mousterian toolmaking were revealed, including flaked blades and points as well as deer, cattle, horse, pig, and fox remains.[71]
1975 Erik Trinkaus' study of Neanderthal feet confirmed they walked like modern humans.
1981 Teeth and part of a jawbone at theBontnewydd Palaeolithic siteby Dr Stephen Aldhouse Green The teeth show evidence of taurodontism, enlarged pulp cavities and short roots, which is characteristic of Neanderthals, and although it is not unique to them it is one of the reasons that the species was identified as Neanderthal 230,000 years BP. nearSt Asaph,Denbighshire,Wales the most north-western fossil site in Eurasia.
1982-1987 "Moshe" skeleton discovered in 1982 by ProfessorOfer Bar-Yosef In 1987 Thermo-luminescence results date Neanderthals at Kebara ( "Moshe" ) to 60,000 BP. Electron spin resonance (ESR) date Es Skhul to 80,000 BP. 60,000 to 80,000 years BP. Kebara Cave,Es Skhul,Israel Neanderthals present in the region from 200,000 to 45,000 BP.
1991 TheTabun Cavecontains a Neanderthal-type female ESR dates showed the Tabun Neanderthal was contemporaneous with modern humans from Skhul and Qafzeh. 120,000 years BP. Nahal Me'arot Nature Reserve,Israel The cave was occupied intermittently during the Lower and Middle Paleolithic ages (500,000 to around 40,000 years ago).
1993 TheScladinaNeanderthal child. The specimen includes a partial cranium and jaw bone, yielding a 127,000 years DNA sample extracted from a tooth molar. He or she is thought to be 8 years and 17 days old at the time of death 100–127,000 years BP. Scladina,Andenne,Belgium Two Neanderthal occupations were identified: one dating from 130,000 years, and the other 40 000 years.
1994 TheSidrón CaveNeanderthal discovery remains of least 12 individuals: three men, three adolescent boys, three women, and three infants inadvertently uncovered. In 2009 Neanderthal ancient mtDNA was partially sequenced in HVR region for three distinct Neanderthals there. Piloñamunicipality,Asturias,northwestern Spain The first sequencing of the Neanderthal Y chromosome was successfully completed from a specimen from Sidrón Cave[72][73]
1997 Matthias Krings et al. are the first to amplify Neanderthal mitochondrial DNA (mtDNA) using a specimen from the Feldhofer grotto in the Neander valley[74] Germany
1997–2000 During new excavations in the Neandertal, additional bone fragments are found, some of which fit the fragments found in 1856, thus pinpointing the exact location of the original find. The exact location had previously been unknown, as the site of the find (the „Kleine Feldhofer Grotte “) was destroyed by limestone mining.[75] Germany
1998 The Lapedo child by archeologist João Zilhão an earlyUpper Paleolithichuman burial in which provided evidence of early modern humans from the west of the Iberian Peninsula. The remains, a largely complete skeleton of an approximately 4-year-old child, buried with pierced shell and red ochre[76] 24,500 years BP. Abrigo do Lagar Velho,Portugal The cranium, mandible, dentition, and postcrania present a mosaic of European early modern human and Neanderthal features.
2000 Neanderthal infant ofMezmaiskaya Caveby Igor Ovchinnikov et al. Ancient DNA was recovered for a mtDNA sequence showing 3.48% divergence from that ofNeanderthal 1,some 2,500 km (1,553 mi) to the west. Phylogenetic analysis places the two in a clade distinct from modern humans, suggesting that their mtDNA types have not contributed to the modern human mtDNA pool.[77] 29,195 ± 965 years B.P.. RussianRepublic of Adygea,Caucasus Mountains
2005 The Max Planck Institute for Evolutionary Anthropology launched the Neanderthal genome project[78] In co-operation with Connecticut-based 454 Life Sciences and the Institute of Quaternary Paleontology and Geology in Zagreb, Croatia. In 2009, the Max Planck Institute announced the "first draft" of a complete Neanderthal genome is completed.[79]
2010 Comparison of Neanderthal genome with modern humans from Africa and Eurasia shows that 1–4% of modern non-African human genome might come from the Neanderthals.
2010 Discovery of Neanderthal tools far away from the influence ofH. sapiensindicate that the species might have been able to create and evolve tools on its own, and therefore be more intelligent than previously thought. Furthermore, it was proposed that the Neanderthals might be more closely related toHomo sapiensthan previously thought and that may in fact be a subspecies of it. Evidence has more recently emerged that these artifacts are probably ofH. sapiens sapiensorigin.[80][81] [[File:|100 px|]]
2012 Charcoal found next to six paintings of seals in the Nerja caves,Malaga,Spain, has been dated to be between 42,300 and 43,500 years old. The paintings themselves have not been dated until 2014 and if their pigment matches the date of the charcoal, they would be the oldest known cave paintings. José Luis Sanchidrián at the University of Cordoba, Spain believes the paintings are more likely to have been painted by Neanderthals than early modern humans.[82][83]
2013 An individual’s jaw, which was unearthed at the Riparo di Mezzena rock-shelter[84] The jawbone has features intermediate between Neanderthals and Homo sapiens suggesting it could be a hybrid. The mitochondrial DNA is Neanderthal. 40,000-30,000 years BP. Riparo di Mezzena, Monti Lessini region, Italy.
2014 Researchers at the University of Colorado Museum in Boulder Neanderthals were not less intelligent than modern humans and "single-factor explanations for the disappearance of the Neandertals are not warranted any more."[85] [[File:|100 px|]]
2014 Prof Thomas Higham of theUniversity of Oxford The most comprehensive dating of Neanderthal bones and tools ever carried out, which demonstrated that Neanderthals died out in Europe between 41,000 and 39,000 years ago.[86][87] [[File:|100 px|]]


Chronology[edit]

Fossils with Neanderthaltraitsinchronologicalorder
Mixed withH. heidelbergensistraits TypicalH. neanderthalensistraits Homo sapienswith some neanderthal-like archaic traits
> 350 ka:Sima de los Huesosc. 500:350kaago[88][89] 135–45 ka:Krapina,Saccopastore skulls,Malarnaud, Altamura, Gánovce,Denisova,Okladnikov,Altai Mountains,Pech de l'Azé,Tabun120–100±5ka,[90]Qafzeh9 100,Shanidar 1 to 980–60 ka,La Ferrassie 170 ka,Kebara60 ka, Régourdou, Mt. Circeo, Combe Grenal,Erd50 ka,La Chapelle-aux Saints 160 ka, Amud I53±8ka,[91][92]Teshik-Tash.
350–200 ka:Pontnewydd225 ka ago. 45–35 ka:Le Moustier45 ka,Feldhofer42 ka, La Quina, l'Horus, Hortus, Kulna, Šipka, Saint Césaire, Bacho Kiro, El Castillo, Bañolas,Arcy-sur-Cure.[93]
200–135 ka:Atapuerca,[94]Vértesszőlős, Ehringsdorf, Casal de'Pazzi, Biache, La Chaise, Montmaurin, Prince, Lazaret, Fontéchevade < 35 ka: Châtelperron, Figueira Brava, Zafarraya 30 ka,[93]Vogelherd 35 to 32 ka[95][96][97]Vindija32,400±80014C B.P.[98](Vi-20831,390±220,Vi-20732,400±1,80014C B.P.),[98]Velika Pećina, < 35Pestera cu Oase35 ka,Mladeč31 ka, Pestera Muierii 30 ka (n/s),[99]Lapedo Child24.5 ka.

Behavior[edit]

The behavioral patterns of the Neanderthals are inferred from their anatomy in combination with the fossil record, the remains and debris they left behind at caves, rock shelters and a number of open air settlements. On the contrary behavior has become the key factor in explaining the variability of the archaeological record.

Neanderthals shared a number of important characteristics with modern humans, such as large brains, manual dexterity and bipedal walking ability, tool use and production, which requires a certain degree of social sophistication. However, Neanderthals lived in relatively small and territorial more isolated groups. Generally small sites and the considerable depth of bone and debris pits suggests that they frequently migrated but retured to the same locations often.[100]

The materials left behind show only minor variations among sites and their foraging systems were considerably less efficient than those of modern human hunter-gatherers, without planned differential use of the landscape necessitating more-muscular limbs and greater endurance.[101][102][103]

Regional differences[edit]

Habitat and culture of "classical Neandertals" since about 125,000 years ago stretched from western Europe all the way to the Levant in the Middle East and beyond the Crimean Peninsula towards Siberia.[104]Behavior was influenced by climate, terrain, seasonal access to drinking water, the presence of game and other food-resources. Migrations patterns were influenced by the search for raw materials for stone tools.


Elmenteitan[edit]

Elmenteitan
Lake Elmenteita satellite image
Alternative namesElmenteitan Culture
Geographical rangeKenya,Africa
PeriodMesolithic,Neolithic
Datesaround 3,300 to 1,300BP
Type siteGamble's Cave
Major sitesGamble's Cave,Gogo Falls,Njoro River Cave
Preceded byKenya Stillbay,Highland Savanna PN
Followed byKenya Wilton

The Elmenteitan culture was a prehistoric lithic industry and pottery tradition with a distinct pattern of land use, hunting and pastoral practice that appeared on the western plains ofKenya,East Africaduring theMesolithic.[105]

Since the lateNeolithiccontinuous food-production, animal domestication and morturary rituals are documented. Progress and growth and continuity is related to wetter periods of increased precipitation, caused by pleistocene fluctuations and retreaing ice sheets.[106][107]


It was named byLouis LeakeyafterLake Elmenteita(also Elementaita), asoda lake,located in theGreat Rift Valley,about 120 km (75 mi) northwest ofNairobi.[108]

Typical object and artifactassemblagesinclude large double-edgedobsidianmicroliths and blades, ceramic bowls and shallow stone vessels. Cattle and small stock was domesticated and and herded in combination with hunting, fishing and foraging. Patterns and degree of subsistence economy varied depending on the location and the local climate.[109][110]Unlike earlier and contemporary cultures, such as the Highland Savanna PN, cremation of the dead took place in caves (Egerton Cave, Keringet Caves). TheNjoro River Cave,first excavated in 1938 byMary Leakeyserved as a mass-burial site. Associated finds include beads, blades, stone bowls, palettes and pottery vessels, that are interpreted as evidence for burial rituals.[111]

The Elmenteitan was first described in 1931 byLouis Leakeywho excavated at the Gamble's Cave and in 1938 at theNjoro River Cave.Leakey had noticed a locally distinct cluster of the lithic industry and a universal pottery tradition in a restricted area on the plains west of the central Great Rift Valley and at theMau Escarpment.[112]Many tool types are present that were common during theAurignacian,but the Elmenteitan is distinguished by the high percentage of long symmetrical two-edgedobsidianblades which were used unmodified and also served as blanks for a great variety of smaller tools.[113]

All Elmenteitan excavation sites are associated with potsherds where pottery was fully developed. Ceramic vessels are mainly undecorated. Several rare, but very distinctive ornamental designs such as irregular punctuations and rim millings were found. Occationally small bowls with outturned rims, handles with holes or horizontal lugs were discovered. [114]


Canadian Arctic Tundra

Auyuittuq National Park,Baffin Island
Area:
1,420,000 km2(550,000 sq mi)
climate:ET
biome:Tundra
elevation: max. 2,500 m (8,200 ft)
coordinates:
N83°4′N74°10′W/ 83.067°N 74.167°W/83.067; -74.167,
S62°22′N79°38′W/ 62.367°N 79.633°W/62.367; -79.633,
E66°37′N61°17′W/ 66.617°N 61.283°W/66.617; -61.283,
W72°2′N125°37′W/ 72.033°N 125.617°W/72.033; -125.617
Geology:

Canadian tundra (light grey area)
Country:
Canada
Protection:

TheCanadian Arctic Tundraoccupies the immediate regions north of thetreelinein Canada's Arctic Lands. These territories encompass 2,600,000 km2(1,000,000 sq mi), 26 % of the country's landmass, including the Arctic Coastal Plains, Arctic Lowlands and theInnuitian Regionof the High Arctic. Tundra accounts for approximately 1,420,000 km2(550,000 sq mi) at theYukon,the [[Northwest Territories]],Nunavut,northeasternManitoba,northernOntario, northernQuebec,northernLabradorand on 36,500 islands, of whichBaffin Islandwith 422,000 km2(163,000 sq mi) is the largest.[115]


Being part of the globaltundra beltof the northern hemisphere Canada's tundra is characterized by extreme conditions with year roundfrozengrounds, long, cold winters, a very short growing season and precipitation lower than most deserts. The fact of low precipitation requires the classification of the tundra as a (cold) desert, yet permafrost causes even lower degrees of drainage and evaporation and as a result the ground, lakes and glaciers hold large quantities of fresh water.

Harsh environmental and weather conditions, slow growth, poor soils, lacking resources and nutrients, barly any variants of land forms support to highly adapted and migratory life forms only. Biodiversity is one dimensional and only sustainable for brief periods of appropriate conditions. Hundreds of species of flowering plants reproduce by budding and division rather than by interaction with insects. [116][117]


Prolonged periods without sun light, frozen soils and strong winds prevent substantial tree growth.[118]Lichen has developed a unique and alternative adaptation strategy, that does not depend on soils for growth. It is one of the most notable, widespread and enduring organisms of the tundra. This symbiotic life form has algae or cyanobacteria live among the filaments of a fungus and benefit from moisture, elementary nutrients and are protected from the environment. Thephotobiont's (algae) photosynthetic processes in turn generates organic carbon sugars for the fungus.


============================)))))))))


Western imperialism in Asia[edit]

Part ofa serieson
New Imperialism
"The Rhodes Colossus" (1892) by Edward Linley Sambourne
History
Theory
See also

Western imperialism in Asiahas its origins inEuropeantraders seeking unrestricted and independent trade with India and the Southeast Asian spice islands or theEast Indiesafter the Ottoman empire had completely shut the Eurasian trade routes in the eastern Mediterranean by the late 15th century. This theAge of Discovery,and the introduction ofearly modern warfareinto what was then called theFar East.By the early 16th century theAge of Sailgreatly expanded Western European influence and development of theSpice Trade under colonialism.There has been a presence of Western Europeancolonial empiresandimperialismin Asia throughout six centuries ofcolonialism,formally ending with the independence of thePortuguese Empire's last colonyEast Timorin 2002. The empires introduced Western concepts ofnationand themultinational state.This article attempts to outline the consequent development of the Western concept of thenation state.

The thrust of European political power, commerce, and culture in Asia gave rise to growing trade incommodities—a key development in the rise of today's modern worldfree marketeconomy. In the 16th century, the Portuguese broke the (overland) monopoly of the Arabs and Italians of trade between Asia andEuropeby thediscovery of the sea route to Indiaaround the Cape of Good Hope.[119]With the ensuing rise of the rivalDutch East India Company,Portuguese influence in Asia was gradually eclipsed.[nb 1]Dutch forces first established independent bases in the East (most significantlyBatavia,the heavily fortified headquarters of the Dutch East India Company) and then between 1640 and 1660 wrestledMalacca,Ceylon,some southern Indian ports, and the lucrativeJapantrade from the Portuguese. Later, the English and the French established settlements inIndiaand established a trade withChinaand their own acquisitions would gradually surpass those of the Dutch. Following the end of theSeven Years' Warin 1763, the British eliminated French influence in India and established theBritish East India Companyas the most important political force on theIndian Subcontinent.

Before theIndustrial Revolutionin the mid-to-late 19th century, demand for oriental goods such as (porcelain,silk,spicesandtea) remained the driving force behind European imperialism, and (with the important exception of British East India Company rule in India) the Western European stake in Asia remained confined largely to trading stations and strategic outposts necessary to protect trade. Industrialisation, however, dramatically increased European demand for Asian raw materials; and the severeLong Depressionof the 1870s provoked a scramble for new markets for European industrial products and financial services in Africa, the Americas, Eastern Europe, and especially in Asia. This scramble coincided with a new era in globalcolonialexpansion known as "theNew Imperialism",which saw a shift in focus from trade andindirect ruleto formal colonial control of vast overseas territories ruled as political extensions of their mother countries. Between the 1870s and the beginning ofWorld War Iin 1914, theUnited Kingdom,France,and theNetherlands—the established colonial powers in Asia—added to their empires vast expanses of territory in theMiddle East,the Indian Subcontinent, andSouth East Asia.In the same period, theEmpire of Japan,following theMeiji Restoration;theGerman Empire,following the end of theFranco-Prussian Warin 1871;Tsarist Russia;and theUnited States,following theSpanish–American Warin 1898, quickly emerged as new imperial powers inEast Asiaand in the Pacific Ocean area.

In Asia, World War I andWorld War IIwere played out as struggles among several key imperial powers—conflicts involving the European powers along with Russia and the rising American and Japanese powers. None of the colonial powers, however, possessed the resources to withstand the strains of both world wars and maintain their direct rule in Asia. Although nationalist movements throughout the colonial world led to the political independence of nearly all of the Asia's remaining colonies,decolonisationwas intercepted by theCold War;and South East Asia,South Asia,the Middle East, and East Asia remained embedded in a world economic, financial, and military system in which the great powers compete to extend their influence. However, the rapid post-war economic development of theEast Asian Tigers,India,thePeople's Republic of China,along with the collapse of theSoviet Union,have loosened European and American influence in Asia, generating speculation today about emergence of modernIndiaandChinaaspotential superpowers.

References[edit]

  1. ^[YCDTOTV.com FAQ "31. Which sets were used for YCDTOTV sketches?" - see "The cave" under Miscellaneous sets. Note: do not correct url formatting as per Wikipedia's Blacklist, June 2010]
  2. ^"AO, le dernier Néandertal - site officiel du film".UGC YM.RetrievedJuly 24,2016.
  3. ^Thomas Henry Huxley:On some fossil remains of man.Kapitel 3 in:Evidence as to man's place in nature.D. Appleton and Company, New York 1863
  4. ^abFriedemann Schrenk, Stephanie Müller:Die Neandertaler,S. 16
  5. ^Martin Kuckenberg:Lag Eden im Neandertal? Auf der Suche nach dem frühen Menschen.Econ Verlag, Düsseldorf 1997, S. 51,ISBN3-430-15773-0
  6. ^"L'homme fossile de La Chapelle-aux-Saints - full text - Volume VI (p. 11–172), Volume VII (p. 21–56), Volume VIII (p. 1–70), 1911–1913".Royal College of Surgeons of England.RetrievedJuly 26,2016.
  7. ^"Marcellin Boule - French geologist".Encyclopædia Britannica.RetrievedJuly 26,2016.
  8. ^"Arthur Keith".Royal Anthropological Institute.RetrievedJuly 26,2016.
  9. ^"La Chapelle-Aux-Saints - The old man of La Chapelle - The original reconstruction of the 'Old Man of La Chapelle' by scientist Pierre Marcellin Boule led to the reason why popular culture stereotyped Neanderthals as dim-witted brutes for so many years".Smithsonian Institution.RetrievedJuly 26,2016.
  10. ^"North American Terrestrial Ecoregions—Level III April 2011"(PDF).Commission for Environmental Cooperation.RetrievedDecember 16,2016.
  11. ^Wood, Bernard (31 March 2011).Wiley-Blackwell Encyclopedia of Human Evolution, 2 Volume Set - edited by Bernard Wood.ISBN9781444342475.RetrievedSeptember 26,2016.
  12. ^"Homo sapiens neanderthalensis".National Center for Biotechnology Information, U.S. National Library of Medicine.RetrievedSeptember 26,2016.
  13. ^Bibliography of Fossil Vertebrates 1954-1958 - C.L. Camp, H.J. Allison, and R.H. Nichols - Google Books.Books.google.ca. Retrieved on 2014-05-24.
  14. ^"Neanderthal in ODE".Oxford Dictionaries.
  15. ^""Neanderthal" in Random House Dictionary (US) & Collins Dictionary (UK) ".Dictionary.com.
  16. ^"Neanderthals' Last Stand Is Traced".The New York Times.September 13, 2006.RetrievedSeptember 26,2016.
  17. ^"WHY DID NEANDERTHALS GO EXTINCT? -...debate whether Homo neanderthalensis should be classified as a subspecies of Homo sapiens..."Smithsonian Insider. August 11, 2015.RetrievedSeptember 26,2016.
  18. ^Hublin, J. J. (2009)."The origin of Neandertals".Proceedings of the National Academy of Sciences.106(38): 16022–7.Bibcode:2009PNAS..10616022H.doi:10.1073/pnas.0904119106.JSTOR40485013.PMC2752594.PMID19805257.
  19. ^Harvati, K.; Frost, S.R.; McNulty, K.P. (2004)."Neanderthal taxonomy reconsidered: implications of 3D primate models of intra- and interspecific differences".Proc. Natl. Acad. Sci. U.S.A.101(5): 1147–52.doi:10.1073/pnas.0308085100.PMC337021.PMID14745010.
  20. ^"Scientists Identify Neanderthal Genes in Modern Human DNA".Sci-News.com. January 30, 2014.RetrievedSeptember 26,2016.
  21. ^Liu, Hua; et al. (2006)."A Geographically Explicit Genetic Model of Worldwide Human-Settlement History".The American Journal of Human Genetics.79(2): 230–237.doi:10.1086/505436.PMC1559480.PMID16826514.Currently available genetic and archaeological evidence is generally interpreted as supportive of a recent single origin of modern humans in East Africa. However, this is wherethe near consensuson human settlement history ends, and considerable uncertainty clouds any more detailed aspect of human colonization history.
  22. ^Currat, M.; Excoffier, L. (November 30, 2004)."Modern Humans Did Not Admix with Neanderthals during Their Range Expansion into Europe -...morphology developed in Europe during the last 350,000 y under the effect of selection and genetic drift..."PLOS Biology.2(12): e421.doi:10.1371/journal.pbio.0020421.PMC532389.PMID15562317.{{cite journal}}:CS1 maint: unflagged free DOI (link)
  23. ^"Neanderthal ANTHROPOLOGY -...Neanderthals inhabited Eurasia from the Atlantic regions..."Encyclopaedia Britannica. January 29, 2015.RetrievedSeptember 26,2016.
  24. ^Shaw, Ian; Jameson, Robert, eds. (1999).A Dictionary of Archaeology.Blackwell. p. 408.ISBN0-631-17423-0.Retrieved1 August2016.
  25. ^"Homo neanderthalensis -...The Mousterian stone tool industry of Neanderthals is characterized by..."Smithsonian Institution. September 22, 2016.RetrievedSeptember 26,2016.
  26. ^Finlayson, C; Pacheco, FG; Rodríguez-Vidal, J; Fa, DA; Gutierrez López, JM; Santiago Pérez, A; Finlayson, G; Allue, E; Baena Preysler, J; Cáceres, I; Carrión, JS; Fernández Jalvo, Y; Gleed-Owen, CP; Jimenez Espejo, FJ; López, P; López Sáez, JA; Riquelme Cantal, JA; Sánchez Marco, A; Guzman, FG; Brown, K; Fuentes, N; Valarino, CA; Villalpando, A; Stringer, CB; Martinez Ruiz, F; Sakamoto, T (October 2006). "Late survival of Neanderthals at the southernmost extreme of Europe".Nature.443(7113): 850–3.Bibcode:2006Natur.443..850F.doi:10.1038/nature05195.ISSN0028-0836.PMID16971951.S2CID4411186.
  27. ^"Homo neanderthalensis Brief Summary".EOL.RetrievedSeptember 26,2016.
  28. ^Peresani, M.; Dallatorre, S.; Astuti, P.; Dal Colle, M.; Ziggiotti, S.; Peretto, C. (2014)."Symbolic or utilitarian? Juggling interpretations of Neanderthal behavior: new inferences from the study of engraved stone surfaces".Journal of Anthropological Sciences = Rivista di Antropologia: Jass.92(92): 233–255.doi:10.4436/JASS.92007.PMID25020018.RetrievedJuly 26,2016.
  29. ^"A high-quality Neandertal genome sequence".Max Planck Institute for Evolutionary Anthropology.RetrievedSeptember 26,2016.
  30. ^Complete Neanderthal genome sequenced: DNA signatures found in present-day Europeans and Asians, but not in Africans,...showed that Neanderthal DNA is 99.7 percent identical to present-day human DNAScienceDaily
  31. ^Colin P.T. Baillie;University of California, Berkeley."Neandertals: Unique from Humans, or Uniquely Human?"(PDF).berkeley.edu.
  32. ^Smithsonian InstitutionNational Museum of American History."Ancient DNA and Neanderthals".si.edu.
  33. ^Sánchez-Quinto, Federico; Botigué, Laura R.; Civit, Sergi; Arenas, Conxita; Ávila-Arcos, María C.; Bustamante, Carlos D.; Comas, David; Lalueza-Fox, Carles (October 17, 2012)."North African Populations Carry the Signature of Admixture with Neandertals".PLOS ONE.7(10): e47765.doi:10.1371/journal.pone.0047765.PMC3474783.PMID23082212.
  34. ^Cite error:The named referenceBBCwas invoked but never defined (see thehelp page).
  35. ^Fu, Qiaomei; Li, Heng; Moorjani, Priya; Jay, Flora; Slepchenko, Sergey M.; Bondarev, Aleksei A.; Johnson, Philip L. F.; Aximu-Petri, Ayinuer; Prüfer, Kay; De Filippo, Cesare; Meyer, Matthias; Zwyns, Nicolas; Salazar-García, Domingo C.; Kuzmin, Yaroslav V.; Keates, Susan G.; Kosintsev, Pavel A.; Razhev, Dmitry I.; Richards, Michael P.; Peristov, Nikolai V.; Lachmann, Michael; Douka, Katerina; Higham, Thomas F. G.; Slatkin, Montgomery; Hublin, Jean-Jacques; Reich, David; Kelso, Janet; Viola, T. Bence; Pääbo, Svante (23 October 2014)."Genome sequence of a 45,000-year-old modern human from western Siberia".Nature.514(7523): 445–449.doi:10.1038/nature13810.hdl:10550/42071.PMC4753769.PMID25341783.
  36. ^Brahic, Catherine."Humanity's forgotten return to Africa revealed in DNA",The New Scientist(February 3, 2014).
  37. ^Wilford, John Noble (December 16, 2013)."Neanderthals and the Dead".New York Times.RetrievedDecember 17,2013.
  38. ^Jaubert, Jacques; Verheyden, Sophie; Genty, Dominique; Soulier, Michel; Cheng, Hai; Blamart, Dominique; Burlet, Christian; Camus, Hubert; Delaby, Serge; Deldicque, Damien; Edwards, R. Lawrence; Ferrier, Catherine; Lacrampe-Cuyaubère, François; Lévêque, François; Maksud, Frédéric; Mora, Pascal; Muth, Xavier; Régnier, Édouard; Rouzaud, Jean-Noël; Santos, Frédéric (2 June 2016) [online 25 May 2016]."Early Neanderthal Constructions deep in Bruniquel Cave in Southwestern France".Nature.534(7605): 111–114.doi:10.1038/nature18291.ISSN0028-0836.PMID27251286.
  39. ^"The Reputed Fossil Man of the Neanderthal. by Professor Wiiliam King"(PDF).Bone and Stone.RetrievedSeptember 26,2016.
  40. ^William King:On the Neanderthal Skull, or Reasons for believing it to belong to the Clydian Period and to a species different from that represented by Man.In:British Association for the Advancement of Science, Notices and Abstracts for 1863, Part II,London, 1864, p. 81 f.
  41. ^William King:The Neanderthal Skull.In:The Anthropological Review.Vol. 1, Nr. 3, 1863, S. 393–394,full text (PDF)
  42. ^Kunzig, Robert."The Year in Science: Human Origins 1997",Discover (magazine)(January 1, 1998) reprinted inContemporary Readings in Physical Anthropology,p. 145 (Alan Almquist ed., Prentice Hall, 2000)
  43. ^"Engis 2".Smithsonian Institution.RetrievedSeptember 27,2015.
  44. ^Philippe-Charles Schmerling:Recherches sur les ossements fossiles découverts dans les cavernes de la Province de Liège.P.-J. Collardin, Liège 1833, pp. 1–66
  45. ^"Engis 3 - individuum Engis 2 - Public NESPOS Space".NESPOS Society.RetrievedSeptember 27,2015.
  46. ^Thomas Henry Huxley:On some fossil remains of man.chapter 3 in:Evidence as to man's place in nature.D. Appleton and Company, New York 1863
  47. ^Thomas Henry Huxley:On some fossil remains of man.Kapitel 3 in:Evidence as to man's place in nature.D. Appleton and Company, New York 1863
  48. ^"Homo neanderthalensis".Patrick Johnson.RetrievedSeptember 26,2016.
  49. ^"Hominids and hybrids: The place of Neanderthals in human evolution by Ian Tattersall and Jeffrey H. Schwartz".National Academy of Sciences.RetrievedSeptember 26,2016.
  50. ^Ian Tattersall:Neandertaler. Der Streit um unsere Ahnen.Birkhäuser Verlag, Basel 1999, S. 81,ISBN3-7643-6051-8.
  51. ^"Calcified remains of Altamura Man yield oldest ever Neanderthal DNA sample".Ancient Origins. April 4, 2015.RetrievedSeptember 3,2016.
  52. ^"'Pit of bones' catches Neanderthal evolution in the act ".Nature. June 19, 2014.RetrievedSeptember 27,2016.
  53. ^"Oldest Ancient-Human DNA Details Dawn of Neandertals".SCIENTIFIC AMERICAN. March 14, 2016.RetrievedSeptember 26,2016.
  54. ^"Homo heidelbergensis- Comparison of Neanderthal and modern human DNA suggests that the two lineages diverged from a common ancestor, most likelyHomo heidelbergensis".Smithsonian Institution.RetrievedSeptember 26,2016.
  55. ^"Oldest ancient-human DNA details dawn of Neanderthals".Nature. March 14, 2016.RetrievedSeptember 27,2016.
  56. ^"Homo antecessor: Common Ancestor of Humans and Neanderthals?".Smithsonian. November 26, 2012.RetrievedSeptember 27,2016.
  57. ^"Le bras d'un pré-Néandertalien trouvé au bord de la Seine] »".Le Monde. October 9, 2014.RetrievedSeptember 28,2016.
  58. ^Faivre, Jean-Philippe; Maureille, Bruno; Bayle, Priscilla; Crevecoeur, Isabelle; Duval, Mathieu; Grün, Rainer; Bemilli, Céline; Bonilauri, Stéphanie; Coutard, Sylvie; Bessou, Maryelle; Limondin-Lozouet, Nicole; Cottard, Antoine; Deshayes, Thierry; Douillard, Aurélie; Henaff, Xavier; Pautret-Homerville, Caroline; Kinsley, Les; Trinkaus, Erik (October 8, 2014)."Middle Pleistocene Human Remains from Tourville-la-Rivière (Normandy, France) and Their Archaeological Context".PLOS ONE.9(10): e104111.doi:10.1371/journal.pone.0104111.PMC4189787.PMID25295956.
  59. ^Tattersall, Ian; Schwartz, Jeffrey H. (1999)."Hominids and hybrids: The place of Neanderthals in human evolution".Proceedings of the National Academy of Sciences.96(13): 7117–9.Bibcode:1999PNAS...96.7117T.doi:10.1073/pnas.96.13.7117.JSTOR48019.PMC33580.PMID10377375.
  60. ^Duarte, Cidália; Mauricio, João; Pettitt, Paul B.; Souto, Pedro; Trinkaus, Erik; Van Der Plicht, Hans; Zilhao, João (1999)."The early Upper Paleolithic human skeleton from the Abrigo do Lagar Velho (Portugal) and modern human emergence in Iberia".Proceedings of the National Academy of Sciences.96(13): 7604–9.Bibcode:1999PNAS...96.7604D.doi:10.1073/pnas.96.13.7604.JSTOR48106.PMC22133.PMID10377462.
  61. ^"L'homme fossile de La Chapelle-aux-Saints - full text - Volume VI (p. 11–172), Volume VII (p. 21–56), Volume VIII (p. 1–70), 1911–1913".Royal College of Surgeons of England.RetrievedJuly 26,2016.
  62. ^"Marcellin Boule - French geologist".Encyclopædia Britannica.RetrievedJuly 26,2016.
  63. ^"Arthur Keith".Royal Anthropological Institute.RetrievedJuly 26,2016.
  64. ^"La Chapelle-Aux-Saints - The old man of La Chapelle - The original reconstruction of the 'Old Man of La Chapelle' by scientist Pierre Marcellin Boule led to the reason why popular culture stereotyped Neanderthals as dim-witted brutes for so many years".Smithsonian Institution.RetrievedJuly 26,2016.
  65. ^"Our Neandertal Brethren: Why They Were Not a Separate Species".SCIENTIFIC AMERICAN. August 1, 2010.RetrievedJuly 26,2016.
  66. ^Wolpoff, MH;Hawks, J;Caspari, R (2000)."Multiregional, not multiple origins"(PDF).American Journal of Physical Anthropology.112(1): 129–36.doi:10.1002/(SICI)1096-8644(200005)112:1<129::AID-AJPA11>3.0.CO;2-K.hdl:2027.42/34270.PMID10766948.
  67. ^Hedges SB (December 2000). "Human evolution. A start for population genomics".Nature.408(6813): 652–3.doi:10.1038/35047193.PMID11130051.S2CID3240904.
  68. ^"Modern humans, Neanderthals shared earth for 1,000 years".ABC News (Australia).1 September 2005. Archived fromthe originalon January 15, 2006.Retrieved19 September2006.
  69. ^"New Evidence On The Role Of Climate In Neanderthal Extinction - The bones of more than 400 Neanderthals have been found since..."Science Daily. September 13, 2007.RetrievedOctober 12,2016.
  70. ^"Neanderthal - Homo neanderthalensis - Details - Key Fossils - La Chapelle-aux-Saints".Encyclopedia of Life.RetrievedJuly 18,2016.
  71. ^"Amud".encyclopaedia britannica.RetrievedOctober 13,2016.
  72. ^"Universitat Pompeu Fabra, Barcelona, A paleogenetical study determines the blood group of Neanderthal man"ArchivedNovember 22, 2012, at theWayback Machine
  73. ^Lalueza-Fox, Carles; Rosas, Antonio; Rasilla, Marco de la (2012)."Palaeogenetic research at the El Sidrón Neanderthal site".Annals of Anatomy - Anatomischer Anzeiger.194(1): 133–137.doi:10.1016/j.aanat.2011.01.014.PMID21482084.
  74. ^Krings, M; Stone, A; Schmitz, RW; Krainitzki, H; Stoneking, M; Pääbo, S (1997). "Neandertal DNA sequences and the origin of modern humans".Cell.90(1): 19–30.doi:10.1016/S0092-8674(00)80310-4.ISSN0092-8674.PMID9230299.S2CID13581775.
  75. ^Schmitz, Ralf W; et al. (2002)."The Neandertal type site revisited: Interdisciplinary investigations of skeletal remains from the Neander Valley, Germany".Proceedings of the National Academy of Sciences.99(20): 13342–13347.doi:10.1073/pnas.192464099.PMC130635.PMID12232049.
  76. ^Duarte, C; Maurício, J; Pettitt, PB; Souto, P; Trinkaus, E; Van Der Plicht, H; Zilhão, J; et al. (1999)."The early Upper Paleolithic human skeleton from the Abrigo do Lagar Velho (Portugal) and modern human emergence in the Iberian Peninsula".Proceedings of the National Academy of Sciences of the United States of America.96(13).PNAS:7604–7609.Bibcode:1999PNAS...96.7604D.doi:10.1073/pnas.96.13.7604.PMC22133.PMID10377462.
  77. ^Ovchinnikov, IV; Götherström, A; Romanova, GP; Kharitonov, VM; Lidén, K; Goodwin, W (2000). "Molecular analysis of Neanderthal DNA from the northern Caucasus".Nature.404(6777): 490–3.doi:10.1038/35006625.PMID10761915.S2CID3101375.
  78. ^"Analysis of one million base pairs of Neanderthal DNA".Nature. November 16, 2006.RetrievedJuly 18,2016.
  79. ^Morgan, James (12 February 2009)."Neanderthals 'distinct from us'".BBC News.Retrieved22 May2009.
  80. ^"Neanderthals More Intelligent Than Thought".September 22, 2010. Archived fromthe originalon September 25, 2010.RetrievedSeptember 27,2010.
  81. ^Benazzi, S.; Douka, K.; Fornai, C.; Bauer, C. C.; Kullmer, O.; Svoboda, J. Í.; Pap, I.; Mallegni, F.; Bayle, P.; Coquerelle, M.; Condemi, S.; Ronchitelli, A.; Harvati, K.; Weber, G. W. (2011). "Early dispersal of modern humans in Europe and implications for Neanderthal behaviour".Nature.479(7374): 525–528.doi:10.1038/Nature10617.PMID22048311.S2CID205226924.
  82. ^Fergal MacErlean (10 February 2012)."First Neanderthal cave paintings discovered in Spain".New Scientist.Retrieved10 February2012.
  83. ^"Our Artistic Relatives: Did Neanderthals Have the Cognitive Capacity to Produce Parietal Art? The Case of El Castillo Cave…".Global Multimedia Protocols Group. March 29, 2014.RetrievedOctober 13,2016.
  84. ^Viegas, Jennifer (27 March 2013)."First Love Child of Human, Neanderthal Found".Discovery News.Retrieved11 April2013.
  85. ^"Neanderthals were not less intelligent than modern humans, scientists find".TheGuardian.com.April 30, 2014.RetrievedApril 30,2014.
  86. ^Cite error:The named referencebbc.co.ukwas invoked but never defined (see thehelp page).
  87. ^Cartmill, Matt; Smith, Fred H. (30 March 2009).The Human Lineage by Matt Cartmill, Fred H. Smith.ISBN9780471214915.RetrievedSeptember 27,2015.
  88. ^Bischoff, J; Shamp, Donald D.; Aramburu, Arantza; Arsuaga, Juan Luis; Carbonell, Eudald; Bermudez De Castro, J.M. (2003). "The Sima de los Huesos Hominids Date to Beyond U/Th Equilibrium (>350kyr) and Perhaps to 400–500kyr: New Radiometric Dates".Journal of Archaeological Science.30(3): 275–280.doi:10.1006/jasc.2002.0834.
  89. ^Arsuaga JL, Martínez I, Gracia A, Lorenzo C (1997). "The Sima de los Huesos crania (Sierra de Atapuerca, Spain). A comparative study".Journal of Human Evolution.33(2–3): 219–81.doi:10.1006/jhev.1997.0133.PMID9300343.
  90. ^Mcdermott, F; Grün, R; Stringer, Cb; Hawkesworth, Cj (May 1993). "Mass-spectrometric U-series dates for Israeli Neanderthal/early modern hominid sites".Nature.363(6426): 252–5.Bibcode:1993Natur.363..252M.doi:10.1038/363252a0.ISSN0028-0836.PMID8387643.S2CID4234812.
  91. ^Rink, W. Jack; Schwarcz, H.P.; Lee, H.K.; Rees-Jones, J.; Rabinovich, R.; Hovers, E. (August 2002). "Electron spin resonance (ESR) and thermal ionization mass spectrometric (TIMS) 230Th/234U dating of teeth in Middle Paleolithic layers at Amud Cave, Israel".Geoarchaeology.16(6): 701–717.doi:10.1002/gea.1017.S2CID129034553.
  92. ^Valladas, Hélène; Merciera, N.; Frogeta, L.; Hoversb, E.; Joronc, J.L.; Kimbeld, W.H.; Rak, Y. (March 1999). "TL Dates for the Neanderthal Site of the Amud Cave, Israel".Journal of Archaeological Science.26(3): 259–268.doi:10.1006/jasc.1998.0334.
  93. ^abRincon, Paul (13 September 2006)."Neanderthals' 'last rock refuge'".BBC News.Retrieved18 May2009.
  94. ^Kreger, C. David."Homo neanderthalensis".ArchaeologyInfo.com.Retrieved16 May2009.
  95. ^"Hohlenstein - Hohlenstein 4".NESPOS Space.RetrievedJuly 24,2016.
  96. ^"Lonetal Sites, including Neanderthal and Middle Paleolithic sites in the Swabian Alb near the city of Ulm - View of Vogelherd about 35 000 years BP...The stratigraphy reveals deposits from [...] Mousterian..."Don's Maps.RetrievedJuly 24,2016.
  97. ^Conard, Nj; Grootes, Pm; Smith, Fh (July 2004). "Unexpectedly recent dates for human remains from Vogelherd".Nature.430(6996): 198–201.Bibcode:2004Natur.430..198C.doi:10.1038/nature02690.ISSN0028-0836.PMID15241412.S2CID4412769.
  98. ^abHigham T, Ramsey CB, Karavanić I, Smith FH, Trinkaus E (January 2006)."Revised direct radiocarbon dating of the Vindija G1 Upper Paleolithic Neandertals".Proceedings of the National Academy of Sciences.103(3): 553–7.Bibcode:2006PNAS..103..553H.doi:10.1073/pnas.0510005103.PMC1334669.PMID16407102.
  99. ^Hayes, Jacqui (2 November 2006)."Humans and Neanderthals interbred".Cosmos.Archived fromthe originalon February 22, 2009.Retrieved17 May2009.
  100. ^"NEANDERTHAL BEHAVIOUR".encyclopaedia britannica.RetrievedOctober 13,2016.
  101. ^"Neanderthal Behavior".Nature.RetrievedOctober 13,2016.
  102. ^"Neanderthals: Facts About Our Extinct Human Relatives".Live Science. April 13, 2016.RetrievedOctober 13,2016.
  103. ^"Rethinking Neanderthals".Smithsonian. April 13, 2016.RetrievedOctober 13,2016.
  104. ^Roura, Eudald Carbonell I. (13 March 2012).High Resolution Archaeology and Neanderthal Behavior: Time and Space in...edited by Eudald Carbonell i Roura.ISBN9789400739222.RetrievedOctober 13,2016.
  105. ^Robertshaw, Peter (1988)."The Elmenteitan: An Early Food-Producing Culture in East Africa".World Archaeology.20(1). Jstor: 57–69.doi:10.1080/00438243.1988.9980056.JSTOR124525.RetrievedDecember 6,2016.
  106. ^"East African Pastoralism: Routes, Outcomes, Questions"(PDF).Alexandria Archive.RetrievedDecember 6,2016.
  107. ^"East African Pastoralism: Routes, Outcomes, Questions"(PDF).Alexandria Archive.RetrievedDecember 6,2016.
  108. ^"Definition of Elmenteitan".Merriam Webster Dictionary.RetrievedDecember 6,2016.
  109. ^"Elmenteitan".Archaeology Wordsmith.RetrievedDecember 6,2016.
  110. ^"A contribution to the study of subsistence patterns of elmenteitan populations with reference to animal bones from Gogo falls in Sourh Nyanza District, Kenya".University of Nairobi Digital Archive.RetrievedDecember 8,2016.
  111. ^"Archaeological Sites Njoro River Cave".Act For Libraries.RetrievedDecember 6,2016.
  112. ^Ehret, Christopher; Posnansky, Merrick (January 1982).The Archaeological and Linguistic Reconstruction of African History by Christopher Ehret, Merrick Posnansky.ISBN9780520045934.RetrievedDecember 8,2016.
  113. ^"The Archaeology of Africa: Food, Metals and Towns y Bassey Andah, Alex Okpoko, Thurstan Shaw".RetrievedDecember 8,2016.
  114. ^"NOTES ON THE STONE AGE CULTURES OF EAST AFRICA. By L. S. B. LEAKEY,Ph.D., F.S.A."(PDF).Biodiversity Library.RetrievedDecember 8,2016.
  115. ^"Physiographic Regions".The Canadian Encyclopedia.RetrievedDecember 9,2016.
  116. ^"The tundra biome".California Academy of Sciences.RetrievedDecember 9,2016.
  117. ^"Canadian Low Arctic Tundra - About the Area".World Wide Fund For Nature.RetrievedDecember 9,2016.
  118. ^"Tundra Biome: Climate, Locations, Plants & Animals".Study.com.RetrievedDecember 9,2016.
  119. ^M. Weisner-Hanks, Early Modern Europe 1450–1789 (Cambridge, 2006)
  120. ^Roberts, Edmund(Digitized 12 October 2007) [First published in 1837].Embassy to the Eastern courts of Cochin-China, Siam, and Muscat: in the U. S. sloop-of-war Peacock... during the years 1832-3-4.Harper & brothers. image 173, p.166.OCLC12212199.{{cite book}}:Check date values in:|date=(help)


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