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Andean condor

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"Vultur" redirects here. For other birds, seeVulture.

Andean condor
Temporal range:2.6–0Ma
↓
LatePliocene–Holocene
Male Andean condor at theCincinnati Zoo.
Female at Doué-la-Fontaine Zoo, France
CITESAppendix I(CITES)[2]
Scientific classificationEdit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Cathartiformes
Family: Cathartidae
Genus: Vultur
Linnaeus,1758
Species:
V. gryphus
Binomial name
Vultur gryphus
Yellow – approximate range/distribution
Synonyms
  • Vultur magellanicusShaw, 1792
  • Sarcoramphus cunturDuméril, 1806
  • Vultur condorShaw, 1809
  • Sarcorhamphus aequatorialisSharpe, 1874
  • Vultur fossilisMoreno& Mercerat, 1891
  • Vultur patruusLönnberg,1902
  • Vultur pratruusEmslie, 1988 (lapsus)

TheAndean condor(Vultur gryphus) is a South AmericanNew World vultureand is the only member of thegenusVultur.It is found in theAndesmountains and adjacent Pacific coasts of western South America. With a maximum wingspan of 3.3 m (10 ft 10 in) and weight of 15 kg (33 lb), the Andean condor is one of the largest flying birds in the world, and is generally considered to be the largestbird of preyin the world.

It is a large black vulture with a ruff of white feathers surrounding the base of the neck and, especially in the male, large white patches on the wings. The head and neck are nearly featherless, and are a dull red color, which may flush and therefore change color in response to the bird's emotional state. In the male, there is a wattle on the neck and a large, dark redcombor caruncle on the crown of the head. The female condor is smaller than the male, an exception to the usualsexual dimorphismseen inbirds of prey.

The condor is primarily ascavenger,feeding oncarrion.It prefers large carcasses, such as those ofdeerorcattle.It reaches sexual maturity at five or six years of age andnestsat elevations of up to 5,000 m (16,000 ft), generally on inaccessible rock ledges. One or two eggs are usually laid. It is one of the world's longest-living birds, with a lifespan of over 70 years in some cases.

The Andean condor is a national symbol ofBolivia,Chile,Colombia,Ecuador,andPeruand plays an important role in thefolkloreandmythologyof the Andean regions. The Andean condor is consideredvulnerableby theIUCN.It is threatened by habitat loss and bysecondary poisoningfrom lead in carcasses killed by hunters.Captive breedingprograms have been instituted in several countries.

Taxonomy and systematics

[edit]
Andean condor in Peru
There is a dark red caruncle (or comb) on the top of the head of the adult male.

The Andean condor was described bySwedishscientistCarl Linnaeusin 1758 in thetenth editionof hisSystema Naturaeand retains its originalbinomialname ofVultur gryphus.[3]The Andean condor is sometimes called the Argentinean condor, Bolivian condor, Chilean condor, Colombian condor, Ecuadorian condor, or Peruvian condor after one of the nations to which it is native. The generic termVulturis directly taken from theLatinvulturorvoltur,which means "vulture".[4]Its specific epithet is derived from a variant of theGreekword γρυπός (grupós,"hook-nosed" ).[5]The word condor itself is derived from theQuechuakuntur.[6][7]

The exacttaxonomicplacement of the Andean condor and the remaining six species ofNew World vulturesremains unclear.[8]Although both are similar in appearance and have similarecological roles,the New World andOld World vulturesevolved from different ancestors in different parts of the world and are not closely related. Just how different the two families are is currently under debate, with some earlier authorities suggesting that the New World vultures are more closely related tostorks.[9]More recent authorities maintain their overall position in the orderAccipitriformesalong with the Old World vultures[10]or place them in their own order, Cathartiformes.[11]TheSouth American Classification Committeehas removed the New World vultures fromCiconiiformesand instead described them asincertae sedis,but notes that a move to Falconiformes or Cathartiformes is possible.[8]

The Andean condor is the only accepted living species of its genus,Vultur.[12]Unlike theCalifornia condor(Gymnogyps californianus), which is known from extensivefossilremains and some additional ones ofcongeners,the fossil record of the Andean condor recovered to date is scant. PresumedPlio-Pleistocenespecies of South American condors were later recognized to be not different from the present species, although one known only from a few rather small bones found in a Pliocene deposit ofTarija Department,Bolivia, may have been a smallerpalaeo subspecies,V. gryphus patruus.[13]

Description

[edit]
Adult male (a) and adult female (b). Note the difference in skin and iris colour, as well as the male's comb.
Necks of dominant males may flush a brilliant yellow colour when feeding at carcasses to signify dominance.

The overall length of the Andean condor can range from 100–130 cm (3 ft 3 in – 4 ft 3 in).[14]Among standard measurements, thewing chordis 75.7–85.2 cm (29.8–33.5 in), thetailis 33–38 cm (13–15 in) and thetarsusis 11.5–12.5 cm (4.5–4.9 in). Measurements are usually taken from specimens reared in captivity.[15]The mean weight is 11.3 kg (25 lb), with the males averaging about a kilogram more at 12.5 kg (28 lb), the females a kilogram less at 10.1 kg (22 lb). Condors possess the heaviest average weight for any living flying bird or animal, ahead oftrumpeter swans(Cygnus buccinator) andDalmatian pelicans(Pelecanus crispus).[16][17]However, other sources claim a mean species body mass of 10.3 kg (23 lb) for the Andean condor.[18]The Andean condor is the largest living land bird capable of flight if measured in terms of average weight and wingspan, although malebustardsof the largest species (far more sexually dimorphic in size) can weigh more.[14][19][20]The mean wingspan is around 283 cm (9 ft 3 in) and the wings have the largest surface area of any extant bird.[20]It has a maximum wingspan of 3.3 m (10 ft 10 in).[21]Among living bird species, only thegreat albatrossesand the two largest species ofpelicanexceed the Andean condor in average and maximal wingspan.[20][22]

Andean condor talons

The adultplumageis all black, except for a frill of white feathers at the base of the neck and, especially in the male, large white bands on the wings, which only appear after the bird's firstmoult.[23]The head and neck, kept meticulously clean, are red to blackish-red, and have few feathers.[24]Their baldness means the skin is more exposed to thesterilizingeffects ofdehydrationand high-altitudeUV light.[25]The crown of the head is flattened, and (in the male) is topped by a dark red comb (also called a caruncle); the skin hanging from its neck is called a wattle.[23]Males also have yellower skin. When condors are agitated (for example, during courtship), their head and neck flush, a clear signal to animals nearby. This flush of colour is especially intense in dominant males when feeding at carcasses, and can happen in just a few seconds.[26]Juveniles are grayish-brown, but with a blackish head and neck, and a brown ruff.[27]

The middle toe is greatly elongated, and the hind one is only slightly developed, while the talons of all the toes are comparatively straight and blunt. The feet are thus more adapted to walking, and are of little use as weapons or organs ofprehensionas in birds of prey andOld World vultures.[28]The beak is hooked, and adapted to tear rotting meat.[29]Theirisesof the male are brown, while those of the female are deep red.[30]They have no eyelashes.[31]Unlike the case with most otherbirds of prey,[32]the female is smaller.[33]

Observation of wing color patterns, and the size and shape of the male's crest, are the best ways of identifying individual Andean condors. Sighting-resighting methods assess the size and structure of populations.[34]

Distribution and habitat

[edit]
Andean condor, in Chilean national parkTorres del Paine

The Andean condor is found inSouth Americain theAndesand theSanta Marta Mountains.In the north, its range begins inVenezuelaand Colombia, where it is extremely rare,[35]then continues south along the Andes in Ecuador, Peru, and Chile, through Bolivia and western Argentina to theTierra del Fuego.[27]In the early 19th century, the Andean condor bred from western Venezuela to Tierra del Fuego, along the entire chain of the Andes, but its range has been greatly reduced due to human activity.[36]Its habitat is mainly composed of open grasslands and alpine areas up to 5,000 m (16,000 ft) in elevation. It prefers relatively open, non-forested areas which allow it to spot carrion from the air, such as thepáramoor rocky, mountainous areas in general.[37]It occasionally ranges to lowlands in eastern Bolivia, northern Peru, and southwestern Brazil,[9]descends to lowland desert areas in Chile and Peru, and is found oversouthern-beechforests inPatagonia.[35]In southern Patagonia, meadows are important for Andean condors as this habitat is likely to have herbivores present. In this region, Andean condor distributions are therefore influenced by the locations of meadows as well as cliffs for nesting and roosting.[38]

Ecology and behavior

[edit]
Andean condors often spend much of their time soaring on mountain updrafts.

The condor soars with its wings held horizontally and itsprimary feathersbent upwards at the tips.[23]The lack of a largesternumto anchor its correspondingly large flight muscles physiologically identifies it as primarily being a soarer. It flaps its wings on rising from the ground, but after attaining a moderate elevation it flaps its wings very rarely, relying on thermals to stay aloft.[39]InThe Voyage of the Beagle,Charles Darwinmentioned watching condors for half an hour without once observing a flap of their wings.[40]It prefers to roost on high places from which it can launch without major wing-flapping effort. Andean condors are often seen soaring near rock cliffs, using the heatthermalsto aid them in rising in the air.[41]Flight recorders have shown that "75% of the birds' flapping was associated with take-off", and that it "flaps its wings just 1% of the time during flight".[42]The proportion of time for flapping is more for short flights. Flapping between two thermal glides is more than flapping between two slope glides.[43]

Like otherNew World vultures,the Andean condor has the unusual habit ofurohidrosis:it often empties itscloacaonto its legs and feet. A cooling effect throughevaporationhas been proposed as a reason for this behavior, but it does not make any sense in the cold Andean habitat of the bird.[10]Because of this habit, their legs are often streaked with a white buildup ofuric acid.[28]

There is a well-developed social structure within large groups of condors, with competition to determine a 'pecking order' by body language, competitive play behavior, andvocalizations.[44]Generally, mature males tend to be at the top of the pecking order, with post-dispersal immature males tending to be near the bottom.[15]

Breeding

[edit]
Egg, CollectionMuseum Wiesbaden

Sexual maturity and breeding behavior do not appear in the Andean condor until the bird is five or six years of age.[45]It may live to 50 years or more, and it mates for life.[46]During courtship displays, the skin of the male's neck flushes, changing from dull red to bright yellow, and inflates.[47]He approaches the female with neck outstretched, revealing the inflated neck and the chest patch, while hissing,[48]then extends his wings and stands erect while clicking his tongue.[30]Other courtship rituals include hissing and clucking while hopping with wings partially spread, and dancing.[25]

A juvenile condor inColca Canyon,Peru.

The Andean condor prefers to roost and breed at elevations of 3,000 to 5,000 m (9,800 to 16,400 ft).[49]Its nest, which consists of a few sticks placed around the eggs, is created on inaccessible ledges of rock. However, in coastal areas of Peru, where there are few cliffs, some nests are simply partially shaded crannies scraped out against boulders on slopes.[36]It deposits one bluish-whiteegg,weighing about 280 g (9.9 oz) and ranging from 75 to 100 mm (3.0 to 3.9 in) in length. Breeding occurs about every second year, in the southern Andes around October, in the central and northern Andes it can be throughout the year.[50]The egg hatches after 54 to 58 days ofincubationby both parents.[30]If the chick or egg is lost or removed, another egg is laid to take its place. Researchers and breeders take advantage of this behavior to double the reproductive rate by taking the first egg away for hand-rearing, causing the parents to lay a second egg, which they are generally allowed to raise.[51]The young are covered with a grayish down until they are almost as large as their parents. They are able to fly after six months,[23]but continue to roost and hunt with their parents until age two, when they are displaced by a new clutch.[52]

Feeding

[edit]

The Andean condor is ascavenger,feeding mainly oncarrion.[39]Wild condors inhabit large territories, often traveling more than 200 km (120 mi) a day in search of carrion.[25]In inland areas, they prefer large carcasses. Naturally, they feed on the largest carcasses available, which can includellamas(Lama glama),alpacas(Lama pacos),rheas(Rhea ssp.),guanacos(Lama guanicoe),deerandarmadillos.Wild individuals could acquire extra carotenoids from vegetal matter contained in carcass viscera and fresh vegetation.[53][54]However, most inland condors now live largely off of domestic animals, which are now more widespread in South America, such ascattle(Bos taurus),horses(Equus caballus),donkeys(Equus asinus),mules,sheep(Ovis aries),domestic pigs(Sus domesticus),domestic goats(Capra hircus) anddogs(Canis familiaris). They also feed on the carcasses of introduced game species such aswild boar(Sus scrofa),rabbits(Oryctolagus cuniculus),foxes(Vulpes vulpes) andred deer(Cervus elaphus). For condors who live around the coast, the diet consists mainly of beached carcasses of marine mammals, largelycetaceans.[30][55]They will also raid the nests of smaller birds to feed on the eggs.[56]Andean condors have been observed to do some hunting of small, live animals, such asrodents,birdsandrabbits,which (given their lack of powerful, grasping feet or developed hunting technique) they usually kill by jabbing repeatedly with their bill.[55]

Andean condor skeleton (Museum of Osteology)

Coastal areas provide a constant food supply, and in particularly plentiful areas, some Andean condors limit their foraging area to several kilometers of beach-front land.[36]They locate carrion by spotting it or by following other scavengers, such ascorvidsor other vultures.[57]It may follow New World vultures of the genusCathartes—theturkey vulture(C. aura), thelesser yellow-headed vulture(C. burrovianus), and thegreater yellow-headed vulture(C. melambrotus)—to carcasses. TheCathartesvultures forage by smell, detecting the scent ofethyl mercaptan,a gas produced by the beginnings of decay in dead animals. These smaller vultures cannot rip through the tougher hides of these larger animals with the efficiency of the larger condor, and their interactions are often an example ofmutual dependencebetween species.[58]However, studies have indicated that Andean condors are fairly proficient at searching out carrion without needing to rely on other scavengers to guide them to it.[59]Black vultures(Coragyps atratus) and several mammalian carnivorous scavengers such asfoxesmay sometimes trackCathartesvultures for carcasses or compete with condors over available carrion but the condor is invariably dominant among the scavengers in its range.[36]A study inPatagoniafound surprisingly that condors were driving the ecology ofpuma(Puma concolor) in the area, apparently by routinely commandeering the powerful cat's kills (often the day following the puma's nighttime kills). It is projected that the condors were able to engage in harassment of the pumas despite the large cat's size and power, and has apparently driven the pumas to increase their kill rate in order to accommodate for their frequent losses to the scavengers.[60][61]Andean condors are intermittent eaters in the wild, often going for a few days without eating, then gorging themselves on several pounds at once, sometimes to the point of being unable to lift off the ground. Because its feet and talons are not adapted to grasping, it must feed while on the ground.[25]Like other carrion-feeders, it plays an important role in itsecosystemby disposing of carrion which would otherwise be a breeding ground for disease.[62]

Andean condors can efficiently absorb a wide variety ofcarotenoidpigments from the vegetal matter within theviscerathat they consume from carcasses. These include carotenoids such asβ-caroteneandechineone.The pigments result in the yellow skin colouration of adult males and their ability to flush them a brilliant yellow during contests for dominance, as well as the colour of the iris and bright orange tongues of both sexes. Captive Andean condors have a lower concentration of carotenoid pigments in their bodies than wild condors, likely because the diet of captive condors is usually restricted to just flesh. An analysis of the droppings of wild condors found that 90% contained vegetal remains, and of those that contained vegetal remains, 35% of them were composed of primarily vegetal matter (around 80% by volume).[26]

Longevity

[edit]

Being a slowly-maturing bird with no known natural predators in adulthood, an Andean condor is a long-lived bird. Longevity and mortality rates are not known to have been extensively studied in the wild. Some estimations of lifespans of wild birds has exceeded 50 years. In 1983, theGuinness Book of World Recordsconsidered the longest-lived bird of any species with a confirmed lifespan was an Andean condor that died after surviving 72 years in captivity, having been captured from the wild as a juvenile of undetermined age.[20]Several species ofparrothave been reported to live for perhaps over 100 years in captivity, but these (at least in 1983) were not considered authenticated.[20]Another early captive-held specimen of condor reportedly lived for 71 years.[20]However, these lifespans have been exceeded by a male, nicknamed "Thaao", that was kept atBeardsley ZooinConnecticut.Thaao was born in captivity in 1930 and died on January 26, 2010, making him 79 years of age.[63]This is the greatest verified age ever known for a bird.[20]

Relationship with humans

[edit]

Conservation status

[edit]
Adult male atTaronga Zoo,Australia
Adult male atMoscow Zoo,Russia

The Andean condor is consideredvulnerableby theIUCNand the PeruvianConservation Organization.[1]As a result of research on its plight, its status was changed to Vulnerable from Near Threatened in 2020, and only about 10,000 individuals remain. It was first placed on the United StatesEndangered Specieslist in 1970,[64]a status which is assigned to an animal that is in danger of extinction throughout all or a significant portion of its range.[65]Threats to its population include loss of habitat needed for foraging, secondary poisoning from animals killed by hunters and persecution.[21]It is threatened mainly in the northern area of its range, and is extremely rare in Venezuela and Colombia, where it has undergone considerable declines in recent years.[66]Because it is adapted to very low mortality and has correspondingly low reproductive rates, it is extremely vulnerable to human persecution,[35]most of which stems from the fact that it is perceived as a threat by farmers due to alleged attacks on livestock.[46]Education programs have been implemented by conservationists to dispel this misconception.[67]Reintroduction programs using captive-bred Andean condors, which release birds hatched in North American zoos into the wild to bolster populations,[67]have been introduced in Argentina, Venezuela, and Colombia. The first captive-bred Andean condors were released into the wild in 1989.[68]When raising condors, human contact is minimal; chicks are fed with glove puppets which resemble adult Andean condors in order to prevent the chicks fromimprintingon humans, which would endanger them upon release as they would not be wary of humans.[69]The condors are kept in aviaries for three months prior to release, where they acclimatize to an environment similar to that which they will be released in.[69]Released condors are tracked by satellite in order to observe their movements and to monitor whether they are still alive.[29]

In response to the capture of all the wild individuals of the California condor, in 1988 theUS Fish and Wildlife Servicebegan a reintroduction experiment involving the release of captive Andean condors into the wild inCalifornia.Only females were released to prevent it becoming aninvasive species.The experiment was a success, and all the Andean condors were recaptured and re-released in South America before the reintroduction of the California condors took place.[70]

In June 2014, local authorities of theAncasmarcaregion rescued two Andean condors that were caged and displayed in a local market as an attraction for tourists.[71]

Role in culture

[edit]
Chancaytextile with flying condors, 1200-1400 AD

The Andean condor is a national symbol of Argentina, Bolivia, Chile, Colombia, Ecuador, Peru andVenezuelan Andesstates. It is thenational birdof Bolivia, Chile, Colombia, and Ecuador.[72]It plays an important role in thefolkloreandmythologyof theSouth AmericanAndean regions,[46]and has been represented in Andean art fromc. 2500 BCEonward,[73]and they are a part of indigenous Andean religions.[74]In Andean mythology, the Andean condor was associated with the sun deity,[75]and was believed to be the ruler of the upper world.[76]The Andean condor is considered a symbol of power and health by many Andean cultures, and it was believed that the bones and organs of the Andean condor possessed medicinal powers, sometimes leading to the hunting and killing of condors to obtain its bones and organs.[29][77]In some versions of Peruvian bullfighting ( "Yawar Fiesta" or "Blood Festival" ), a condor is tied to the back of a bull, where it pecks at the animal as bullfighters fight it. The condor generally survives and is set free.[78]

The Andean condor is a popular figure onstampsin many countries, appearing on one for Ecuador in 1958, Argentina in 1960, Peru in 1973, Bolivia in 1985, Colombia in 1992, Chile in 1935 and 2001, andVenezuelain 2004.[79]It has also appeared on the coins and banknotes of Colombia and Chile.[80]

National coat of arms
Others coat of arms
Condor in popular culture
  • Hunting for condor. Chile, 1854.[81]
    Hunting for condor. Chile, 1854.[81]
  • Argentinian gauchos lassoing a condor, 1895.
    Argentiniangauchoslassoing a condor, 1895.
  • Chilean 60-centavo stamp, 1935.
    Chilean 60-centavo stamp, 1935.

References

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  1. ^abBirdLife International (2020)."Vultur gryphus".IUCN Red List of Threatened Species.2020:e.T22697641A181325230.doi:10.2305/IUCN.UK.2020-3.RLTS.T22697641A181325230.en.Retrieved19 November2021.
  2. ^"Appendices | CITES".cites.org.Retrieved2022-01-14.
  3. ^Linnaeus, C(1758).Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata(in Latin). Vol. v.1. Holmiae (Stockholm): Laurentii Salvii. p. 86.V. maximus, carúncula verticali longitudine capitis.
  4. ^Simpson, D.P. (1979).Cassell's Latin Dictionary(5th ed.). London: Cassell Ltd. p. 883.ISBN0-304-52257-0.
  5. ^Liddell, Henry George; Robert Scott (1980).Greek-English Lexicon, Abridged Edition.Oxford: Oxford University Press.ISBN0-19-910207-4.
  6. ^J. Simpson; E. Weiner, eds. (1989). "Raven".Oxford English Dictionary(2nd ed.). Oxford: Clarendon Press.ISBN0-19-861186-2.
  7. ^"A Quechua metaphor for a plane: Kuntur-man =" looking like a Condor "".Quechua.org.uk. Archived fromthe originalon 2014-06-06.Retrieved2010-03-20.
  8. ^abRemsen, J. V. Jr.; Cadena, C. D.; Jaramillo, A.; Nores, M.; Pacheco, J. F.; Robbins, M. B.; Schulenberg, T. S.; Stiles, F. G.; Stotz, D. F. and Zimmer, K. J. (2007).A classification of the bird species of South America.Archived2009-03-02 at theWayback MachineSouth American Classification Committee. Retrieved on 2007-10-15
  9. ^abSibley, Charles Gald; Monroe, Burt Leavelle (1990).Distribution and Taxonomy of Birds of the World.Yale University Press.ISBN978-0-300-04969-5.[permanent dead link]
  10. ^abSibley, Charles G.; Jon E. Ahlquist (1991).Phylogeny and Classification of Birds: A Study in Molecular Evolution.Yale University Press.ISBN0-300-04085-7.
  11. ^Ericson, P. G.P; Anderson, C. L; Britton, T.; Elzanowski, A.; Johansson, U. S; Kallersjo, M.; Ohlson, J. I; Parsons, T. J; et al. (2006)."Diversification of Neoaves: Integration of molecular sequence data and fossils".Biology Letters.2(4): 543–7.doi:10.1098/rsbl.2006.0523.PMC1834003.PMID17148284.
  12. ^"Vultur gryphus".Integrated Taxonomic Information System. Archived fromthe originalon 2008-12-09.Retrieved2008-01-13.
  13. ^Fisher, Harvey L. (1944)."The skulls of the Cathartid vultures"(PDF).Condor.46(6): 272–296.doi:10.2307/1364013.JSTOR1364013.Archived fromthe original(PDF)on 2015-11-17.Retrieved2013-04-05.
  14. ^abdel Hoyo, J; Elliot, A; Sargatal, J (1996).Handbook of the Birds of the World.Vol. 3.Barcelona:Lynx Edicions.ISBN84-87334-20-2.
  15. ^abFerguson-Lees, James; Christie, David A. (2001).Raptors of the World.Boston: Houghton Mifflin.ISBN0-618-12762-3.
  16. ^Dunning, John B. Jr., ed. (2008).CRC Handbook of Avian Body Masses(2nd ed.). CRC Press.ISBN978-1-4200-6444-5.
  17. ^Wallace, M. P.; Temple, S. A. (1987). "Competitive interactions within and between species in a guild of avian scavengers".The Auk.104(2): 290–295.doi:10.1093/auk/104.2.290.
  18. ^Atanasov, A. T. (2007). "The near to linear allometric relationship between total metabolic energy per life span and body mass of nonpasserine birds".Bulgarian Journal of Veterinary Medicine.10(4): 235–245.
  19. ^"Andean CondorVultur gryphus".birdlife.org.Birdlife International.
  20. ^abcdefgWood, Gerald (1983).The Guinness Book of Animal Facts and Feats.Enfield, Middlesex, England: Guinness Superlatives.ISBN978-0-85112-235-9.
  21. ^abReading, Richard P.; Miller, Brian (2000)."Andean Condor".Endangered Animals: A Reference Guide to Conflicting Issues.Greenwood Press. p.16.ISBN0-313-30816-0.
  22. ^Harrison, Peter (1991).Seabirds: An Identification Guide.Houghton Mifflin Harcourt.ISBN978-0-395-60291-1.
  23. ^abcdHilty, Stephen L. (1977).A Guide to the Birds of Colombia.Princeton University Press. p. 88.ISBN0-691-08372-X.
  24. ^"Behavior of the Andean Condor".clemetzoo.com.Cleveland Metroparks Zoo. Archived fromthe originalon 19 December 2006.Retrieved2007-01-10.
  25. ^abcdLutz, Dick; Lutz, Richard L. (2002).Patagonia: At the Bottom of the World.DIMI Press. pp.71–74.ISBN0-931625-38-6.
  26. ^abBlanco, Guillermo; Hornero-Méndez, Dámaso; Lambertucci, Sergio A.; Bautista, Luis M.; Wiemeyer, Guillermo; Sanchez-Zapata, José A.; Garrido-Fernández, Juan; Hiraldo, Fernando; Donázar, José A. (2013-06-13)."Need and Seek for Dietary Micronutrients: Endogenous Regulation, External Signalling and Food Sources of Carotenoids in New World Vultures".PLOS ONE.8(6): e65562.doi:10.1371/journal.pone.0065562.ISSN1932-6203.PMC3681859.PMID23785435.
  27. ^abBlake, Emmet Reid (1953).Birds of Mexico: A Guide for Field Identification.University of Chicago Press. pp.262–263.ISBN0-226-05641-4.
  28. ^abFeduccia, J. Alan (1999).The Origin and Evolution of Birds.Yale University Press. p. 300.ISBN0-226-05641-4.
  29. ^abc"Andean Condor".Zoological Society of San Diego.Retrieved2007-01-10.
  30. ^abcdFriends of the Zoo."Andean Condor".Smithsonian National Zoological Park. Archived fromthe originalon 2007-10-09.Retrieved2008-01-08.
  31. ^Fisher, Harvey L. (1942)."The Pterylosis of the Andean Condor".Condor.44(1): 30–32.doi:10.2307/1364195.JSTOR1364195.
  32. ^Andersson, Malte B. (1994).Sexual selection.Princeton University Press. p. 269.ISBN978-0-691-00057-2.Retrieved19 July2010.
  33. ^Arnold, Caroline; Wallace, Michael Phillip; Wallace, Michael (1993).On the brink of extinction: the California condor.Houghton Mifflin Harcourt. p. 11.ISBN978-0-15-257990-6.Retrieved19 July2010.
  34. ^Méndez, D.; Marsden, S.; Lloyd, H. (2019)."Assessing population size and structure for Andean Condor Vultur gryphus in Bolivia using a photographic 'capture-recapture' method".Ibis.161(4): 867–877.doi:10.1111/ibi.12681.
  35. ^abc"Species factsheet: Vultur Gryphus".BirdLife International. 2004. Archived fromthe originalon 2006-03-05.Retrieved2008-01-04.
  36. ^abcdHaemig, PD (2007)."Ecology of Condors".Ecology Online Sweden. Archived fromthe originalon 4 December 2007.RetrievedMarch 30,2009.
  37. ^"Habitat of the Andean Condor".Cleveland Metroparks Zoo. Archived fromthe originalon 20 December 2006.Retrieved2007-01-10.
  38. ^Pérez-García, J.M.; Sánchez-Zapata, J.A.; Lambertucci, S.A.; Hiraldo, F.; Donázar, J.A (2018). "Low-frequency, threatened habitats drive the large-scale distribution of Andean Condors in southern Patagonia".Ibis.160(3): 647–658.doi:10.1111/ibi.12563.hdl:11336/90584.
  39. ^abWehner, Ross; del Gaudio, Renee; Jankowski, Kazia (2007).Moon Peru.Avalon Travel. p. 180.ISBN978-1-56691-983-8.
  40. ^Darwin, Charles (1909).The Voyage of the Beagle.P.F. Collier. p.201.
  41. ^Benson, Sara; Paul Hellander (2007).Peru.Lonely Planet Publications. p.53.ISBN978-1-74059-749-4.
  42. ^"Andean condor birds 'flap wings just 1% of the time'".BBC. 14 July 2020.Retrieved14 July2020.
  43. ^Williams, H. J.; Shepard, E. L. C.; Holton, Mark D.; Alarcón, P. a. E.; Wilson, R. P.; Lambertucci, S. A. (2020-07-10)."Physical limits of flight performance in the heaviest soaring bird".Proceedings of the National Academy of Sciences.117(30): 17884–17890.Bibcode:2020PNAS..11717884W.doi:10.1073/pnas.1907360117.ISSN0027-8424.PMC7395523.PMID32661147.
  44. ^Donazard, José A; Feijoo, Juan E. (2002)."Social structure of Andean Condor roosts: Influence of sex, age, and season".Condor.104(1). Cooper Ornithological Society: 832–837.doi:10.1650/0010-5422(2002)104[0832:SSOACR]2.0.CO;2.hdl:10261/65935.ISSN0010-5422.S2CID35374983.Archived fromthe originalon 2012-09-04.Retrieved2008-01-10.
  45. ^"Andean Condor (Vultur Gryphus) ".The Peregrine Fund.Archived fromthe originalon 5 February 2007.Retrieved2007-01-10.
  46. ^abcTait, Malcolm (2006).Going, Going, Gone: Animals and Plants on the Brink of Extinction.Sterling Publishing. p.22.ISBN1-84525-027-3.
  47. ^Whitson, Martha A; Whitson, Paul D. (1968)."Breeding Behavior of the Andean Condor (Vultur Gryphus) "(PDF).Condor.71(1). Cooper Ornithological Society: 73–75.doi:10.2307/1366056.JSTOR1366056.Retrieved2007-01-10.
  48. ^Gailey, Janet; Bolwig, Neils (1973)."Observations on the Behavior of the Andean Condor (Vultur Gryphus) "(PDF).Condor.75(1). Cooper Ornithological Society: 60–68.doi:10.2307/1366535.JSTOR1366535.Retrieved2007-01-10.
  49. ^Fjeldså, Jon; Krabbe, Niels (1990).Birds of the High Andes.Apollo Books. p. 90.ISBN87-88757-16-1.
  50. ^Avian Report. "Andean Condor Nesting Habits".https://avianreport.com/andean-condor-nesting/
  51. ^ National Research Council (1992).Scientific Bases for the Preservation of the Hawaiian Crow.National Academies Press. p. 74.ISBN0-309-04775-7.
  52. ^Cisneros-Heredia, Diego F. (2006)."Notes on breeding, behaviour and distribution of some birds in Ecuador".Bulletin of the British Ornithologists' Club126(2): 153–164 contains a record of a juvenile accompanying an adult male in July, too early to have been of that year's cohort.
  53. ^Blanco, G.; Hornero-Méndez, D.; Lambertucci, S.A.; Donázar, J.A.; Bautista, L.M.; Wiemeyer, G.; Sanchez-Zapata, J.A.; Garrido-Fernández, J.; Hiraldo, F. (2013)."Need and seek for dietary micronutrients: endogenous regulation, external signaling and food sources of carotenoids in New World vultures"(PDF).PLOS ONE.8(6): e65562.Bibcode:2013PLoSO...865562B.doi:10.1371/journal.pone.0065562.PMC3681859.PMID23785435.
  54. ^Blanco, G.; Bautista, L.M.; Hornero-Méndez, D.; Lambertucci, S.A.; Wiemeyer, G.; Sanchez-Zapata, J.A.; Hiraldo, F.; Donázar, J.A. (2014)."Allometric deviations of plasma carotenoids in raptors"(PDF).Ibis.156(3): 668–675.doi:10.1111/ibi.12155.hdl:10261/98308.
  55. ^abKidd, Travis."Vultur gryphus (Andean condor)".Animal Diversity Web.Retrieved2023-02-11.
  56. ^"Andean Condor (Vultur Gryphus) ".National Geographic.Archived fromthe originalon June 12, 2007.Retrieved2007-01-10.
  57. ^Snyder, Noel F. R.; Helen Snyder (2006).Raptors of North America: Natural History and Conservation.Voyageur Press. p.45.ISBN0-7603-2582-0.
  58. ^Muller-Schwarze, Dietland (2006).Chemical Ecology of Vertebrates.Cambridge University Press. p. 350.ISBN0-521-36377-2.
  59. ^Lambertucci, S. A., Speziale, K. L., Rogers, T. E., & Morales, J. M. (2009).How do roads affect the habitat use of an assemblage of scavenging raptors?Biodiversity and Conservation, 18(8), 2063-2074.
  60. ^Elbroch, L. M., & Wittmer, H. U. (2013).Nuisance ecology: do scavenging condors exact foraging costs on pumas in Patagonia?PLOS ONE, 8(1), e53595.
  61. ^Perrig, P. L., Donadio, E., Middleton, A. D., & Pauli, J. N. (2017).Puma predation subsidizes an obligate scavenger in the high Andes.Journal of applied ecology, 54(3), 846-853.
  62. ^Gomez, Luis G.; Houston, David C.; Cotton, Peter; Tye, Alan (1994)."The role of greater yellow-headed vultures Cathartes melambrotus as scavengers in neotropical forest".Ibis.136(2): 193–196.doi:10.1111/j.1474-919X.1994.tb01084.x.Archived fromthe originalon 29 January 2008.Retrieved2008-01-06.
  63. ^Zoo Family Mourns Death of Oldest Living Andean Condor in Captivity | Connecticut's Beardsley ZooArchived2012-05-20 at theWayback Machine.Beardsleyzoo.org (2010-01-26). Retrieved on 2012-12-19.
  64. ^"Species Profile: Andean Condor".United States Department of Fish and Wildlife. Archived fromthe originalon March 3, 2009.Retrieved2007-10-16.
  65. ^"Endangered Species Program".United States Department of Fish and Wildlife.Archivedfrom the original on 2007-09-21.Retrieved2007-10-16.
  66. ^Beletsky, Les (2006).Birds of the World.JHU Press. p. 70.ISBN0-8018-8429-2.
  67. ^abRoach, John (2004-07-22)."Peru's Andean Condors Are Rising Tourist Attraction".National Geographic News.National Geographic.Archived fromthe originalon July 26, 2004.Retrieved2007-01-10.
  68. ^Conservation and Research for Endangered Species."Andean Condor Reintroduction Program".Zoological Society of San Diego. Archived fromthe originalon 2006-10-10.Retrieved2007-01-10.
  69. ^abPullin, Andrew S. (2002).Conservation Biology.Cambridge University Press. p. 234.ISBN0-521-64482-8.
  70. ^"California condor, (Gymnogyps californianus)".U.S. Fish and Wildlife Service. Archived fromthe originalon 2013-04-01.Retrieved2007-08-14.
  71. ^(in Spanish)Rescatan dos cóndores andinos que eran exhibidos a turistas en Calca CuscoArchived2017-09-13 at theWayback Machine,Andina.com.pe,12 June 2014
  72. ^MacDonald, Tina; MacDonald, Duncan."National Birds".Archivedfrom the original on 6 October 2007.Retrieved2007-10-06.
  73. ^Werness, Hope B. (2004).The Continuum Encyclopedia of Animal Symbolism in Art.Continuum International Publishing Group. p. 103.ISBN0-8264-1525-3.
  74. ^Howard-Malverde, Rosaleen (1997).Creating Context in Andean Cultures.Oxford University Press. p. 16.ISBN0-19-510914-7.
  75. ^Mundkur, Balaji (1983).The Cult of the Serpent.SUNY Press. p. 129.ISBN0-87395-631-1.
  76. ^Mills, Alice; Parker, Janet; Stanton, Julie (2006).Mythology: Myths, Legends and Fantasies.New Holland Publishers. p. 493.ISBN1-77007-453-8.
  77. ^"History of the Andean Condor".Cleveland Metroparks Zoo. Archived fromthe originalon 19 December 2006.Retrieved2007-01-10.
  78. ^Kokotovic, Misha (2007).The Colonial Divide in Peruvian Narrative:Social Conflict and Transculturation.Sussex Academic Press. p. 49.ISBN978-1-84519-184-9.
  79. ^"Andean Condor".Bird Stamps. Archived from the original on October 23, 2000.Retrieved2008-01-15.{{cite web}}:CS1 maint: unfit URL (link)
  80. ^"A Field Guide to the Birds on Banknotes".Krause Publications.Retrieved2008-01-16.
  81. ^Gay, Claudio (1854)."Image 61 of Atlas of the Physical and Political History of Chile [Plates, Volume 1]".Library of Congress.Paris: E. Thunot.Retrieved2023-02-11.
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