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Haddock

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Haddock
Haddock at theNew England Aquarium
Scientific classificationEdit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Gadiformes
Family: Gadidae
Genus: Melanogrammus
T. N. Gill,1862
Species:
M. aeglefinus
Binomial name
Melanogrammus aeglefinus
Synonyms

Thehaddock(Melanogrammus aeglefinus) is asaltwaterray-finned fishfrom thefamilyGadidae,the truecods.It is the only species in themonotypicgenusMelanogrammus.It is found in the NorthAtlantic Oceanand associated seas, where it is an important species for fisheries, especially in northern Europe, where it is marketed fresh, frozen andsmoked;smoked varieties include theFinnan haddieand theArbroath smokie.Other smoked versions include long boneless, the fileted side of larger haddock smoked in oak chips with the skin left on the fillet.

Description

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The haddock has the elongated, tapering body shape typical of members of the cod family.[2]It has a relatively small mouth which does not extend to below the eye; with the lower profile of the face being straight and the upper profile slightly rounded, this gives its snout a characteristic wedge-shaped profile. The upper jaw projects beyond the lower more so than in theAtlantic cod.[3]There is a rather small barbel on the chin.[4]There are threedorsal fins,the first being triangular in shape[5]and these dorsal fins have 14 to 17 fin rays in the first, 20 to 24 in the second, and 19 to 22 in the third. There are also twoanal finsand in these there are 21 to 25 fin rays in the first and 20 to 24 fin rays in the second.[3]The anal and dorsal fins are all separated from each other.[6]The pelvic fins are small with an elongated first fin ray.[7]

The upper side of the haddock's body varies in colour from dark grey brown to nearly black while the lower part of the body is dull silvery white. It has a distinctive blacklateral linecontrasting with the whitish background colour and which curves slightly over thepectoral fins.It also has a distinctive oval black blotch or ‘thumbprint’, sometimes called the "Devil's thumbprint",[8]which sits between the lateral line and the pectoral fin,[9]a feature which leads to the name of the genusMelanogrammuswhich derives fromGreek"melanos"meaning "black" and"gramma"meaning letter or signal.[4]The dorsal, pectoral, andcaudal finsare dark grey in colour while the anal fins are pale matching the colour of the silvery sides, with black speckles at their bases. The pelvic fins are white with a variable amount of black spots. Occasionally there are differently coloured variants recorded which may be barred, golden on the back or lack the dark shoulder blotch.[3]

The longest haddock recorded was 94 centimetres (37 inches) in length and weighed 11 kilograms (24 pounds). However, haddock are rarely over80 cm (31+12in) in length and the vast majority of haddocks caught in the United Kingdom measure between30 and 70 cm (12 and27+12in).[9]In easternCanadawaters, haddock range in size from 38 to 69 cm (15 to 27 in) in length and 0.9 to 1.8 kg (2 lb 0 oz to 3 lb 15 oz) in weight.[8]

Distribution

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The haddock has populations on either side of the north Atlantic but it is more abundant in the eastern Atlantic than it is on the North American side. In the north-east Atlantic it occurs from theBay of Biscaynorth toSpitzbergen;however, it is most abundant north of theEnglish Channel.It also occurs aroundNovaya Zemlyaand theBarents Seain the Arctic.[6]The largest stocks are in theNorth Sea,off theFaroe Islands,offIcelandand the coast of Norway but these are discrete populations with little interchange between them. Off North America, the haddock is found from westernGreenlandsouth toCape Hatteras,but the main commercially fished stock occurs fromCape Codand theGrand Banks.[9]

Habitat and biology

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Fins,barbelandlateral lineon a haddock. Haddock have threedorsal finsand twoanal fins.

The haddock is ademersal specieswhich occurs at depths from 10 to 450 m (33 to 1,500 ft; 5.5 to 250 fathoms), although it is most frequently recorded at 80 to 200 m (300 to 700 ft; 40 to 100 fathoms). It is found over substrates made up of rock, sand, gravel or shells and it prefers temperatures of between 4 and 10 °C (39 and 50 °F). Off Iceland and in the Barents Sea, haddock undergo extensive migrations, but in the north western Atlantic its movements are more restricted, consisting of movements to and from their spawning areas. They reach sexual maturity at 4 years old in males and 5 years old in females, except for the population in the North Sea which matures at ages of 2 years in males and 3 years in females. The overall sex ratio is roughly 1:1, but in shallower areas, females predominate, while the males show a preference for waters further offshore.[6]

The fecundity of the females varies with size: a fish of 25 cm (10 in) length bears 55,000 eggs while a fish at 91 cm (36 in) has 1,841,000 eggs. Spawning takes place from depths of around 50 to 150 m (200 to 500 ft; 30 to 80 fathoms). In the northwestern Atlantic spawning lasts from January to July, although it does not occur simultaneously in all areas, and in the northeastern Atlantic the spawning season runs from February to June, peaking in March and April.[6]The eggs are pelagic with a diameter of1.2 to 1.7 millimetres (364to9128in), and they take one to three weeks to hatch. Following metamorphosis, the past larval fish remain pelagic until they attain a length of around 7 cm (3 in), when they settle to a demersal habit.[10]Their growth rate shows considerable regional variation and fish at one year old can measure17 to 19 cm (6+12to7+12in), at 2 years old 25 to 36 cm (10 to 14 in), up to75 to 82 cm (29+12to32+12in) at 13 years old. Their lifespan is around 14 years.[6]The most important spawning grounds are in the waters off the central coast ofNorway,off the southwest ofIceland,and over theGeorges Bank.[10][11]The fish which spawn in inshore waters are normally smaller and younger fish than those which occur in offshore areas. The younger fish have a spawning season which is less than half of that of the larger and older stock offshore. Once hatched the larvae do not appear to travel far from their spawning grounds, however some larvae spawning off the west coast ofScotlandare transported into the North Sea through theFair Isle-ShetlandGap or to the northeast of Shetland.[12]

In their larval stages, haddock mainly feed on the immature stages ofcopepodsand the pelagic post-larvae up to 3–10 cm (1–4 in) in length prey onkrill,Larvaceans,decapodlarvae, copepods and on small fish. Once they have reached the settled, demersal, post-larval stage,benthicinvertebrates become increasingly important although they still feed on pelagic organisms such as krill, however the benthic invertebrates form an increasing part of their diet as they grow. Adults prey on fish such assand eels,Trisopterus esmarkii,American plaice,gobies,European sprat,andAtlantic herring,as well ascapelin,silver hake,American eelsandargentines.[4]When a number of fish taken at the same time have their stomach contents sampled, the majority of stomachs contain similar prey, this suggests that haddocks feed in shoals.[10]Shellfish,sea urchins,brittlestarsand worms are also important prey,[9]especially in the winter.[12]Juvenile haddock are an important prey for larger demersal fish, including othergadoids,while seals prey on the larger fish.[10]

The recorded growth rates of haddock underwent significant change over the 30 to 40 years up to 2011.[10]Growth has been more rapid in recent years, with haddock attaining adult size much earlier than was noted 30–40 years ago. However, the degree to which these larger, younger fish contribute to reproductive success of the population is unknown. The growth rates of haddock, however, have slowed in recent years. There is some evidence which indicates that these slower growth rates may be the result of an exceptionally large year class in 2003.[11]The haddock stock periodically has higher than normal productivity; for example in 1962 and 1967, and to a lesser extent, 1974 and 1999. These result in a more southerly distribution of the fish and have a strong effect on the biomass of the spawning stock, but because of high fishing mortality, these revivals do not have any lasting effect on the population. In general, there was above average recruitment from the 1960s up to the early 1980s, similar to recruitment for Atlantic cod andwhiting,this has been called thegadoid outburst.There was strong recruitment in 1999 but since then, the recruitment rate has been very low.[10]

Parasites

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Cod and related species are plagued by parasites. For example, thecod worm,Lernaeocera branchialis,starts life as acopepod,a small, free-swimming crustaceanlarva.The first host used by cod worm is aflatfishorlumpsucker,which they capture with grasping hooks at the front of their bodies. They penetrate the lumpsucker with a thinfilamentwhich they use to suck its blood. The nourished cod worms then mate on the lumpsucker.[13][14]

The female worm, with her now fertilized eggs, then finds a cod, or a cod-like fish such as a haddock orwhiting.There, the worm clings to thegillswhile it metamorphoses into a plump, sinusoidal, wormlike body, with a coiled mass of egg strings at the rear. The front part of the worm's body penetrates the body of the cod until it enters the rear bulb of the host'sheart.There, firmly rooted in the cod's circulatory system, the front part of the parasite develops like the branches of a tree, reaching into the mainartery.In this way, the worm extracts nutrients from the cod's blood, remaining safely tucked beneath the cod's gill cover until it releases a new generation of offspring into the water.[10][13][14]

Taxonomy and etymology

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The haddock was first formallydescribedasGadus aeglefinusin 1758 byCarolus Linnaeusin the 10th edition of volume one of hisSystema naturaewith atype localitygiven as "European seas".[15]In 1862Theodore Nicholas Gillcreated the genusMelanogrammuswithM. aeglefinusas its only species.[16]The 5th edition ofFishes of the Worldclassifies the haddock within thesubfamilyGadinae,the typical cods, of the family Gadidae, which is within thesuperfamilyGadoideaof the order Gadiformes.[17]

The generic nameMelanogrammusmeans "black line", a reference to the black lateral line of this species. The specific name is alatinisationof the vernacular namesegrefinandeglefin,used in France and England.[18]

Fisheries

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Landings of haddock in the eastern Atlantic in the period 1980–2022. Data from ICES.[19][20][21][22][23]

Haddock is fished year-round using gear such as Danishseine nets,trawlers,long linesandgill netsand is often caught in mixed species fishery with othergroundfishspecies such ascodand whiting. The main fishing grounds in the eastern Atlantic are in theBarents Sea,aroundIceland,around theFaeroe Islands,in theNorth Sea,Celtic Sea,and in theEnglish Channel.Landings in the eastern Atlantic have fluctuated around 200–350 thousand tonnes in the period 1980–2017. During the 1980s, the largest portion of the catch was taken at Rockall but from about 2000, the majority of the catch is caught in the Barents Sea. All the stocks in eastern Atlantic are assessed byICES,which publish a recommendations on an annual basis for Total Allowable Catch.[19][20][21][22][23]

Landings of haddock in the western Atlantic in the period 1960–2022[24]

In the western Atlantic the eastern Georges Bank haddock stock is jointly assessed on an annual basis by Canada and the United States and the stock is collaboratively managed through the Canada–United States Transboundary Management Guidance Committee, which was established in 2000. The commercial catch of haddock in North America was approximately 40–60 thousand tonnes per year between 1920 and 1960. This declined sharply in the late 1960s to between 5 and 30 thousand tonnes per year. Despite a few good years post-1970, landings have not returned to historical levels.[11]

Haddock currently resides on the Greenpeace seafood red list due to concerns regarding the impact of bottom trawls on the marine environment.[25]In contrast, Monterey Bay Aquarium considers haddock a "good alternative".[26]Many haddock fisheries have been certified as sustainable by the Marine Stewardship Council.[27]All seven stocks assessed in the eastern Atlantic are currently considered by ICES to be harvested sustainably. The haddock populations in the western Atlantic (offshore grounds ofGeorges BankoffNew EnglandandNova Scotia) are also considered to be harvested sustainably.[28]

As food

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Haddock, roast
Nutritional value per 100 g (3.5 oz)
Energy469 kJ (112 kcal)
0.0 g
Dietary fiber0.0 g
0.93 g
24.24 g
VitaminsQuantity
%DV
Thiamine (B1)
3%
0.040 mg
Riboflavin (B2)
3%
0.045 mg
Niacin (B3)
29%
4.632 mg
Pantothenic acid (B5)
3%
0.150 mg
Vitamin B6
20%
0.346 mg
Folate (B9)
3%
13 μg
Vitamin C
0%
0.00 mg
MineralsQuantity
%DV
Calcium
3%
42 mg
Iron
8%
1.35 mg
Magnesium
12%
50 mg
Phosphorus
19%
241 mg
Potassium
13%
399 mg
Zinc
4%
0.48 mg
Percentages estimated usingUS recommendationsfor adults,[29]except for potassium, which is estimated based on expert recommendation fromthe National Academies.[30]

Haddock is very popular as a food fish. It is sold fresh or preserved by smoking, freezing, drying, or to a small extent canning. Haddock, along withAtlantic codandplaice,is one of the most popular fish used in Britishfish and chips.[31]

Smoked Haddock served withonionsandred peppers

When fresh, the flesh of haddock is clean and white and its cooking is often similar to that of cod.[5]A fresh haddock fillet will be firm and translucent and hold together well but less fresh fillets will become nearly opaque.[32]Young, fresh haddock and cod fillets are often sold asscrodinBoston, Massachusetts;[33]this refers to the size of the fish which have a variety of sizes, i.e., scrod, markets, and cows.[34]Haddock is the predominant fish of choice in Scotland in a fish supper.[35]It is also the main ingredient of Norwegian fishballs (fiskeboller).[36]Unlike cod, haddock is not an appropriate fish forsaltingand preservation is more commonly effected bydryingandsmoking.[37]

The smoking of haddock was highly refined inGrimsby.Traditional Grimsby smoked fish(mainly haddock, but sometimes cod) is produced in the traditional smokehouses in Grimsby, which are mostly family-run businesses that have developed their skills over many generations.[38]Grimsby fish market sources its haddock from the North East Atlantic, principally Iceland, Norway and the Faroe Islands. These fishing grounds are sustainably managed[39]and have not seen the large scale depreciation in fish stocks seen in EU waters.[40]

One popular form of haddock isFinnan haddiewhich is named after the fishing village ofFinnan or Findonin Scotland, where the fish was originally cold-smoked over smoulderingpeat.Finnan haddie is oftenpoachedin milk and served forbreakfast.[41][42]

The town ofArbroathon the east coast of Scotland produces theArbroath smokie.This is a hot-smoked haddock which requires no further cooking before eating.[43]

Smoked haddock is naturally an off-white colour and it is frequentlydyedyellow, as are other smoked fish. Smoked haddock is the essential ingredient in the Anglo-Indian dishkedgeree,[5]and also in the Scottish dishCullen skink,achowder-like soup.[44]

References

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  1. ^Sobel, J. (1996)."Melanogrammus aeglefinus".IUCN Red List of Threatened Species.1996:e.T13045A3406968.doi:10.2305/IUCN.UK.1996.RLTS.T13045A3406968.en.Retrieved12 April2024.
  2. ^Barnes, M.K.S. (2008). Tyler-Walters H.; Hiscock K. (eds.)."Melanogrammus aeglefinusHaddock ".Marine Life Information Network: Biology and Sensitivity Key Information Reviews.Marine Biological Association of the United Kingdom.Retrieved13 April2018.
  3. ^abcHenry B. Bigelow & William C. Schroeder (1953)."HaddockMelanogrammus aeglefinus".Fishes of the Gulf of Maine.United States Fish and Wildlife Service.Retrieved14 April2018.
  4. ^abcFroese, Rainer; Pauly, Daniel (eds.) (2018)."Melanogrammus aeglefinus"inFishBase.February 2018 version.
  5. ^abc"Haddock".British Sea Fishing. 20 August 2012.Retrieved7 January2019.
  6. ^abcdeDaniel M. Cohen; Tadashi Inada; Tomio Iwamoto & Nadia Scialabba, eds. (1990).VOL. 10 Gadiform Fishes of the World (Order Gadiformes) An Annotated and Illustrated Catalogue of Cods, Hakes, Grenadiers and other Gadiform Fishes Known to Date(PDF).Food and Agriculture Organizationof the United Nations. pp. 54–55.ISBN978-92-5-102890-2.Retrieved2 April2018.
  7. ^Alwyne Wheeler(1992).The Pocket Guide to Salt Water Fishes of Britain and Europe(1997 ed.). Parkgate Books. p. 47.ISBN978-1855853645.
  8. ^ab"Haddock".Fisheries and Oceans Canada.Retrieved14 April2018.
  9. ^abcdTorry Research Station."The haddock".Food and Agriculture Organization of the United Nations. Archived fromthe originalon 26 December 2018.Retrieved14 April2018.
  10. ^abcdefg"Haddock"(PDF).ICES Fish Map.International Council for the Exploration of the Sea.Archived fromthe original(PDF)on 15 April 2018.Retrieved14 April2018.
  11. ^abc"NEFSC Ref Doc".NOAA.Retrieved14 April2018.
  12. ^ab"Haddock".Topics Marine & Fisheries.Scottish GovernmentRiaghaltas na h-Alba. Archived fromthe originalon 14 March 2020.Retrieved13 April2018.
  13. ^abMatthews B (1998)An Introduction to ParasitologyPage 73–74. Cambridge University Press.ISBN978-0-521-57691-8.
  14. ^abExtraordinary Animals: An Encyclopedia of Curious and Unusual Animals.Greenwood Press.2007.ISBN9780313339226.
  15. ^Eschmeyer, William N.;Fricke, Ron & van der Laan, Richard (eds.)."Gadus aeglefinus".Catalog of Fishes.California Academy of Sciences.Retrieved6 June2020.
  16. ^Eschmeyer, William N.;Fricke, Ron & van der Laan, Richard (eds.)."Melanogrammus".Catalog of Fishes.California Academy of Sciences.Retrieved6 June2020.
  17. ^Nelson, J.S.;Grande, T.C.; Wilson, M.V.H. (2016).Fishes of the World(5th ed.). Hoboken, NJ:John Wiley & Sons.pp. 299–302.doi:10.1002/9781119174844.ISBN978-1-118-34233-6.LCCN2015037522.OCLC951899884.OL25909650M.
  18. ^Christopher Scharpf; Kenneth J. Lazara (19 April 2020)."Order GADIFORMES: Families EUCLICHTHYIDAE, MORIDAE, MACRURONIDAE, MERLUCCIIDAE, RANICIPITIDAE, BREGMACEROTIDAE, MURAENOLEPIDIDAE and GADIDAE".The ETYFish Project Fish Name Etymology Database.Christopher Scharpf and Kenneth J. Lazara.Retrieved6 June2020.
  19. ^abICES (9 June 2023).Haddock (Melanogrammus aeglefinus) in Division 5.a (Iceland grounds)(Report). ICES Advice: Recurrent Advice.doi:10.17895/ices.advice.21828339.
  20. ^abICES (30 June 2023).Haddock (Melanogrammus aeglefinus) in Division 7.a (Irish Sea)(Report). ICES Advice: Recurrent Advice.doi:10.17895/ices.advice.21840801.
  21. ^abICES (24 November 2023).Haddock (Melanogrammus aeglefinus) in Division 5.b (Faroes grounds)(Report). ICES Advice: Recurrent Advice.doi:10.17895/ices.advice.21907938.
  22. ^abICES (30 June 2023).Haddock (Melanogrammus aeglefinus) in Divisions 7.b-k (southern Celtic Seas and English Channel)(Report). ICES Advice: Recurrent Advice.doi:10.17895/ices.advice.21840807.
  23. ^abICES (30 June 2023).Haddock (Melanogrammus aeglefinus) in Subarea 4, Division 6.a, and Subdivision 20 (North Sea, West of Scotland, Skagerrak)(Report). ICES Advice: Recurrent Advice.doi:10.17895/ices.advice.21840795.
  24. ^FAO (2024)."Global Capture Production. In: Fisheries and Aquaculture. Rome. [Cited Friday, April 26th 2024]".
  25. ^"Greenpeace International".Archived fromthe originalon 4 July 2019.Retrieved4 July2019.
  26. ^"Haddock".Monterey Bay Aquarium.Retrieved15 February2021.
  27. ^"MSC".
  28. ^Kirsten Weir (July 2009)."The Great Haddock Revival".The Scientist.Retrieved14 April2018.
  29. ^United States Food and Drug Administration(2024)."Daily Value on the Nutrition and Supplement Facts Labels".FDA.Archivedfrom the original on 27 March 2024.Retrieved28 March2024.
  30. ^National Academies of Sciences, Engineering, and Medicine; Health and Medicine Division; Food and Nutrition Board; Committee to Review the Dietary Reference Intakes for Sodium and Potassium (2019). Oria, Maria; Harrison, Meghan; Stallings, Virginia A. (eds.).Dietary Reference Intakes for Sodium and Potassium.The National Academies Collection: Reports funded by National Institutes of Health. Washington, DC: National Academies Press (US).ISBN978-0-309-48834-1.PMID30844154.Archivedfrom the original on 9 May 2024.Retrieved21 June2024.
  31. ^"Fish and chips".Seafish Business to Business Website. Archived fromthe originalon 11 October 2008.Retrieved14 April2018.
  32. ^"Products > Whole Fish & Seafood > HaddockMelanogrammus aeglefinus".SevenSeasFoods. Archived fromthe originalon 14 April 2018.Retrieved14 April2018.
  33. ^G. Stephen Jones (23 July 2010)."Cod or Scrod – What's the Difference?".Reluctant Gourmet.Retrieved14 April2018.
  34. ^"Haddock"(PDF).Rastelli Seafood.Retrieved14 April2018.
  35. ^"Traditional Fish and Chips in Batter".Scotland's Enchanted Kingdom.Retrieved14 April2018.
  36. ^Lois Sinaiko Webb & Lindsay Grace Cardella (2011).Holidays of the World Cookbook for Students, 2nd Edition: Updated and Revised.ABC-CLIO. p. 2018.ISBN978-0313383946.
  37. ^"Haddock fish identification, its habitats, characteristics, fishing methods".All Fishing Guide.Retrieved14 April2018.
  38. ^Grimsby Traditional Fish Smokers GroupArchived3 September 2010 at theWayback Machine
  39. ^Icelandic Request on the Evaluation of Icelandic Cod and Haddock Management PlanArchived6 December 2010 at theWayback Machine
  40. ^European Commission, Communication on Fishing Opportunities for 2009. May 2008
  41. ^Full recipe for Finnan Haddie from Scottish chef John QuigleyArchived24 October 2007 at theWayback Machine
  42. ^"Traditional Scottish Recipes - Finnan Haddie".RampantScotland.Retrieved14 April2018.
  43. ^"What Is A 'Smokie?'".Iain R Spink "The Smokie Man". Archived fromthe originalon 30 December 2012.Retrieved14 April2018.
  44. ^"Cullen skink".BBC.Retrieved14 April2018.

Further reading

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